Lecture 28: Forestomach Fermentative Digestion-2 Flashcards
Grazers or browsers: selectively feeding on highly digestible material
Browsers
What do grazers mainly eat
Grass
How do grazers and browsers handle the high concentration of tannins in their diet
Parotid gland produces proline-rich proteins which bind and neutralize tannins
Which has a larger parotid gland, browsers or grazers
Browsers
Browser or grazer: rumen content is not layered
Browser
Describe the rumen structure/contents of the browser
No fiber mat, no stratification allowing fora more homogenous distribution of particles
Browser or grazer: high viscosity fluid, separation due to floatation/sedimentation less possible, gas bubbles evenly distributed
Browser
Browser or grazer: low viscosity fluid: separation due to flotation sedimentation and clear separation of gaseous layer
Grazer
What stimulates rumen papilla growth
VFA’s
VFA concentration is highest where __activity is highest
Microbial
How does the distribution of the ruminal papilla differ in the grazer and browser
Browsers have a more evenly distributed rumen papilla, whereas the papilla are greatest at the fiber mat water boundary in grazers where microbial activity and VFA’s are highest
Where browsers have the most papilla and greatest absorption of VFA in the dorsal, atrium or ventral rumen
Evenly distributed throughout all 3
Where are the most papilla and greatest absorption of VFA’s in the grazer rumen, dorsal, atrium or ventral rumen
Atrium which is boundary between fiber mat and sediment
How does papilla size indicate activity in the rumen
Small papilla indicate decrease microbial activity and inappropriate pH, whereas large papilla indicates increase microbial activity
What epithelium layer are VFA’s absorbed through
Stratified epithelium
How does a long mean retention time affect utilization of fiber
Improves utilization of fiber
How does a long mean retention time affect feed intake
Restricts because the intake of roughage is limited mostly by the capacity of the forestomach
Who has a greater mean retention time of particle passages through the rumen, browsers or grazers
Grazers have greater retention time for food particles through the rumen to have time for microbial fermentation
Who has a longer fluid retention time in the rumen, grazers or browsers
Browsers
Why is it important for grazers to have a higher fluid throughput in the rumen
Water passing through rumen is needed to:
1. Wash out microbial proteins to SI to become available to calf (increase microbial yield)
2. Saliva contains bicarbonate buffer which will neutralize acidic rumen from VFA’s
3. Move sediment waste from the bottom that is taking up space
Why is food particle throughout in rumen faster in browsers
Rely more on cell wall content (carbs and proteins) which is easily fermentable, unlike grazers which need to ferment fiber
Removing waste and microbes stimulates ___growth by maintaining appropriate conditions in the rumen. This increases ___ yield from the forestomach
Bacterial, microbial
Maintaining rumen activity promotes what phase of growth
Log phase
What is quorum sensing
Mechanism by which bacteria through small signal molecules called autoinducers regulate population density so as higher densities bacteria stop reproducing and making microbial protein
What are the 2 functions of the omasum
- Absorption of water, electrolytes, VFA and bicarbonate
- Transport of ingesta to abomasum
What is the absorption of water so important in the omasum
Needs to dilute solution before entering abomasum in order to optimize activity in abomasum and maintain pH2-3
What is the function of omasal laminae
Surface extensions
Who has a greater water reabsorption, grazers or browsers and why
Grazers, because omasum is larger in size
Why does more water flow through grazers vs browsers stomachs
Helps remove waste and microbial protein and facilitates better fiber fermentation
What are the end products of microbial production of VFA’s by breaking down cellulose and starch
- Acetate
- Proprionate
- Butyrate