Long Exam 2 Flashcards
evolved sometime during the Cambrian period, 500 million years ago during the Cambrian explosion, almost at the same time when invertebrates began to develop
Chordates
May have evolved from some freshwater forms as all modern _____ possess glomerular kidneys designed to remove excess water from the body
Chordates
Early fossils of these have been all recovered from marine sediments and even modern protochordate are all marine forms
Chordates
also found in marine forms such as myxinods and sharks
glomerular kidneys
hagfish class Myxini, jawless, boneless, and sightless fish that exhibit unique behavors
myxinods
chordates evolved from them, includes echinoderms, hemichordates, pogonophrans, etc.
deuterostome
fossils of the earliest vertebrates were known from its, about 400 mya
Silurian-Devonian Period
do not have eyes, ears, and jaws but are considered an important organism in studying chordates
Amphioxus
precursor of a backbone, humans carry a vestiges of these (disk in the spine)
Notochord
first mammals with bony jaws
fish
with this happening, organisms become bigger
four fold increase in genes
a group of marine animals that spend most of their lives attached to docks, rocks or the undersides of boats
tunicate
are semi-transparent barrel-shaped marine animals that move through the water by contracting bands of muscles which ring the body
salp
porous walls of its home allow food particles to flow into an inner chamber where this blue, wispy animal feeds
larvacean
showed how reptiles sporting innovations like jaws and legs flourished on land
monitor lizards
origin theory given by Johannes Muller on 1860 based on comparative studies of larval stages of echinoderms and hemichordates
echinoderm origin
resembles echinoderm larvae such as Bipinnaria, Auricularia, Dipleurula, and Doliolaria, which possess ciliary bands and an apical tuft of cilia
Tornaria larva
proposed that echinoderm larvae gave rise to chordates by neoteny
Johannes Muller
W. Garstang
DeBeers
are deuterostomes and possess mesodermal skeletal elements
Echinoderms
fossil echinoderms discovered from the Ordovician period (450 mya)
Calcichordata
Asymmetrical animals that demonstrate affinities with both echinoderms and chordates but their skeleton is made up of CaCO3
calcichordata
Have large pharynx with series of gill slits, each covered with flaps for filter feeding, a small segmented body, and postanal tail
calcichordata
functions for filter-feeding appears to have evolved in diverse groups of animals during the Cambrian-Ordovician periods
perforated pharynx
bones are made up of hydrated calcium and phosphate
vertebrates
suggested by Romer in 1959 wherein ancestral deuterostomes were sedentary tentacle feeders whose mucous-laden ciliated tentacles served to trap planktons as they were waved in water as do the modern lophophorates and pterobranch
hemichordates, Cephalodiscus and Rhabdopleura
hemichordate origin
Pharyngeal slits evolved in this ancestors, which made pharynx sieve-like to trap planktons as the water current passed through it
hemichordate
possess both ciliated arms and pharyngeal gill slits
pretobranch
W. Garstand (1928) and N.J. Berill (1955) – gave importance to the tadpole-like larva or urochordates which carries typical chordate characters, namely, a notochord in tail along with segmented myotomes, dorsal hollow nerve cord, sense organs, and pharyngeal gill slits
Urochordate origin
suggested that chordates evolved from some sessile filter-feeding urochordate by the larval stage evolving into adulthood by neoteny and by losing the sedentary adult stage
Garstang
Chamberlain studies the primitive and advanced characteristics of cephalochordates and proposed that while extant cephalochordates possess all chordate characters in a typical state, they also show some primitive features of non-chordates, such as the absence of heart, head, sense organs, respiratory pigment, filter
-feeding mode of food capture and excretion by solenocytes
Cephalochordate origin
earliest chordate whose fossil have been discovered from Burgess Shale in British Columbia, Canada.
Pikaia gracilens
show a streamlined, ribbon-shaped, 5 cm long body having notochord in the posterior two-thirds of the body and myomeres
Amphioxus
combined all above theories of echinoderm, urochordata, cephalochordate origins and proposed that the common ancestor of echinoderms and chordates was a sessile ciliary arm feeder that lived in the plankton-rich environment of the Cambria
Combined theory of E.J.W Barriton
modern version of these have evolved from similar ancestors by retaining the original mode of feeding
Echinodermata
perforation of the pharynx with gill slits, must have evolved in a large number of groups which have been a much superior method of food gathering by filtering water as compared to ciliated arm feeding
pharyngothremy
marine animals and lack a bony or cartilaginous skeleton; suspension feeders
cephalochordates and urochordates
have an endoskeleton and some are terrestrial and most use jaws to feed
vertebrates
Four basic characteristics of phylum chordata at some time in their history:
notochord
dorsal hollow nerve cord
pharyngeal slits
post anal tail
dorsal supporting rod
notochord
Replaced by a vertebral column in adult vertebrates (anlagen/precursor of the vertebral column)
notochord
in contrast to invertebrates having a ventral solid nerve cord
dorsal hollow nerve cord
have a hollow nerve cord meaning that the cord contains a canal that is filled with fluid
vertebrates
humans have this fluid in their hollow nerve cord
central canal cerebrospinal fluid
persist in adult fishes, but in most vertebrates are only seen during embryological development
pharyngeal slits
Water passes into the mouth and the pharynx going through here, which are supported by gill bars and used for gas exchange
pharyngeal slits
represents a posterior elongation of the boy extending beyond the anus
post anal tail
Extension of the chordate locomotor organ, the segmental musculature, and notochord
post anal tail
study of the development of embryos from fertilization until they become fetuses or at point which you can distinguish the species
embryology
comparison of embryo development across species
comparative embryology
all embryos pass from single cells to multicelled what
zygotes
clumps of cells
morula
hollow balls of cells
blastula
The process of embryogenesis begins with an ____ or ovum being fertilized by a ____ cell to form a zygote
egg
sperm
zygote is surrounded by a strong membrane made up of this, which the sperm has managed to penetrate
glycoproteins
formed 24 hours after the egg and sperm nuclei fuse
zygote
Over the next three days, the zygote undergoes a number of cell divisions, a process referred to as _____
cleavage
process once the embryo has reached 8-celled stage, involves the tight binding of the cells to create a compact shere
compaction
16-celled embryo that the embryo turns after compaction
Morula
cavity that developed in the morula after one day of compaction
Blastocele
structure that formed after the cells inside the blastocele compact and flatten
Blastocyst
moves towards the womb, where it implants itself in the lining after 24 hours
Blastocyst
Over the next week, the mass of cells rapidly divides, giving rise to a disc-shaped structure that has two layers:
animal pole
vegetal pole
layer becomes the embryo and amniotic cavity
animal pole
layer develops into the yolk sac
vegetal pole
occurs where the blood system starts to appear in the placenta and blood cells are produced by the yok sac, a streak of cells becomes apparent on the embryonic disc
Gastrulation
study of development from fertilization to embryo
embryology
Reveals ancestry
embryology
Nature of relationship from ancestor to offspring can be studied
embryology
ontogeny recapitulates phylogeny.”
who said?
Ernst Haeckel:
Recapitulation is not only relationship between embryos and ancestors”
Gavin De Beer
according to De Beers, these may or may not be present from their ancestors or descendants
embryonic structures
Can be lost (vestigial) or retained to adulthood
embryonic structures
may be repeated in earlier stages but not in later stages
developmental stages
may or may not be altered in the descendant
developmental sequences
sex cells, haploid chromosomes that unite to form diploid organism
gametes
Perform meiotic division
gametes
gametes for males, have various head appearance such
sperm
head apperance of sperm (7)
spherical
spatulate
hooked
lancet-shaped
spiraled
capped (acrosome)
used for movement in sperms
tail
part of the sperm that contains digestive enzymes
acrosome
part of the sperm that contains 23 chromosomes
nucleus
part of the sperm cell that contains many mitochondria
collar
causes sperm to swim
flagellum
are made up of microlecithal (small yolk), mesolecithal (moderate yolk), and
macrolecithal (abundant yolk)
egg cell
small yolk in egg cell
microlecithal
moderate yolk in egg cell
mesolecithal
abundant yolk in egg cell
macrolecithal
process of formation of gametes i.e. sperms and
ovary from the primary sex organs in
all sexually reproducing organisms
gametogenesis
plays most significant role in the process of gametogenesis
meiosis
spermatogenesis formula
1:2:4
differentiation of an ovum into a cell competent to further develop when fertilized
oogenesis
developed from the primary oocyte by maturation
ovum
Initiated in the embryonic stage
oogenesis
Primary oocyte in oogenesis count of chromosomes
46/2N
secondary oocyte in oogenesis count of chromosomes
23/1n
oogonium count of chromosomes
46, 2N
ootid count of chromosomes
23, 1N
a cell in ovary which may undergo meiotic division to form an ovum
oocyte
Immature egg cell that will eventually break free from the follicle and travel down the fallopian tube – at which point it’s called an egg or ovum
oocyte
single cell released from either of the female reproductive organs, the ovaries, which is capable of dev
eloping into a new organism when fertilized (united) with a sperm cell
ovum
developed and released by ovarian follicle
oocyte
surrounds and nourishes the egg
vitelline membrane
in therian mammals, egg is enclosed in this along with corona radiata, cells of ovarian follicle
zona pellucida
present in amphioxus; eutherian mammals (egg yolk)
microlecithal
of, relating to, or belonging to the Eutheria, a subclass of mammals all of which have a placenta and reach an advanced state of development before birth.
eutherian
egg yolk present in lampreys; some fishes; amphibians
mesolecithal
egg yolk present in most fishes; reptiles; birds; monotremes (egg-laying mammals)
macrolecithal
Egg is enclosed after ovulation in jelly layer in what group of animals
amphibians
Egg is enclosed after ovulation in albumen in what group of animals
birds
Egg is enclosed after ovulation in horny, membranous in what animals
fishes
Egg is enclosed after ovulation in calcareous shells in what group of animals
reptiles, birds
Most animals proceed through these stages during development (5)
zygote
early cleavage stages
gastrulation
segmentation (inc. neurulation)
organogenesis
establish polarity and body axes
blasutla
where germ layers are established
gastrulation
this involves neurulation
segmentation
in therian mammals, penetration of sperm from corona radiata to vitelline membrane of the egg
fertilzation
Involves enzymatic and physical interactions between sperm acrosome and egg cortex
fertilization
when this happens the diploid # of chromosomes are restored
union of gametes
hollow sphere of cells produced during the development of an embryo by repeated cleavage of a fertilized egg
vertebrate blastula
epithelial covering layer in blastula
blastoderm
fluid-filled cavity of the blastula
blastocoel
daughter cells
blastomeres
Composed of single tissue layer with hundred of cells
blastula
nutrition of developing embryo; process of cleaving & blastula is dependent on the yolk present
yolk
multiple numbered cells ready for uterine implantation
morula
Cleavage can be:
▪
2-celled
▪
4-celled
▪
8-celled
▪
16-celled
▪
32-celled stages
epiblast or epimere; developing embryo nourished by the vegetal pole (dorsal cells – ectoderm)
animal pole
hypoblast or hypomere, developing yolk, nourishes the embryo (ventral cells – endoderm)
vegetal pole
” (total cleavage furrows penetrate the entire yolk; equal sized blastomeres)
“holoblastic
characterized by or being incomplete cleavage as a result of the presence of an impeding mass of yolk material
meroblastic
holoblastic types (4)
radial
bilateral
spiral
rotational
meroblastic types (2)
discoidal
superficial
holoblastic; unequal-sized blastomeres in amphibians
mesolecithal
larger blastomeres; nourishes the embryo; slower development
vegetal pole
towards the animal pole
blastocoel
“meroblastic” (partial cleavage); unequal sized blastomeres in birds
macrolecithal
large size yolk mass; too great to be penetrated by cleavage furrow in birds
vegetal pole
relatively small (blastoderm)
animal pole
have a microlecithal; holoblastic; unequal sized blastomeres
mammals
structure formed in the early development of mammals
blastocyst
possesses an inner cell mass (ICM) which subsequently forms the embryo
blastocyst
outer layer of the blastocyts containing cells
trophoblast
surrounded by the trophoblast, a fluid-filled cavity
blastocoel
– trophoblast gives rise to this
placenta
Greek word means “a sprout”
blastos
Greek word means “bladder, capsule)
kystis
formation of three germ layers
gastrulation
epiblast or epimere, developing embryo
animal pole
cells migrate to the interior of the blastula, consequently forming two (in diploblastic animals) or into three (tripoblastic animals
gastrulation
embryo during gastrulation is called
gastrula
infolding of cell sheet into embryo
invagination
inturning of cell sheet over the basal surface of an outer layer
involution
migration of individual cells into the mesoderm
ingresion
splitting or migration of one sheet into two sheets
delamination
expansion of one cell sheet over other cell sheets
epiboly
3 germ layers
ectoderm
mesoderm
endoderm
– from epiblast (animal pole) outer layer
ectoderm
middle layer; mesenchyme
mesoderm
from hypoblast (vegetal pole), innermost layer
endoderm
mesolgea is present in what (diploblastic or tripoblastic)
diploblastic
instead of mesoglea, ___ is present in triploblastic animals
mesoderm
central nervous system, retina and lens, cranial and sensory, ganglia and nerves, pigment cells (melanocytes), head connective tissue, epidermis of skin, hair, mammary glands
from what germ layer?
ectoderm
musculoskeletal system, circulatory system, dermis of skin, connective tissue, urogenital system, heart, blood (lymph cells), and spleen
what layer
mesoderm
gastrointestinal system (gut); stomach, colon, liver, pancreas, urinary bladder, lining of urethra, epithelial parts of trachea, lungs, pharynx, thyroid, parathyroid, intestine
what layer
endoderm
formed where cells are entering the embryo
blastopore
two major group of animals can be distinguished according to the blastopore’s fate:
deuterostome
protostome
anus forms from the blastopore
deuterostome
blastopores develops into mouth
protostome
folds inward toward animal pole, double-walled cup
vegetal pole
body cavity (coelom) in gastrulation
gastrocoel
uppermost layer of cells
ectoderm
primitive gut (yolk) lined by endoderm
archenteron
middle layer formed from dorsolateral outpocketing of archenteron
mesoderm
anlagen of nervous system (primitive), formed from dorsal wall of archenteron
notochord
anlagen of nervous system (primitive), formed from dorsal wall of archenteron
notochord
roof of archenteron, forms the notochord in the midline and somites (series of
paired mesodermal tissue blocks)
chordamesoderm
gut tube. Pouch form
schizocoel
formed by splitting of the hypoblast in the somites
coelom
indirectly filled with maternal fluid and enlarges
blastocoel
flattened inner cell mass forms primitive streak (notochord -> CNS)
blastoderm
forms extraembryonic membranes (fetal membranes) and body of embryo (musculoskeletal and circulatory system); mesenchymal tissue
mesoderm
forms by schizocoel (pouch form)
coelom
inner cell mass + primitive streak
embryoblast
notochord (anlagen of CNS); promotes neurulation
primitive streak
overlaps with gastrulation establishing the central nervous system (CNS)
neurulation
coelom in neurulation
neurocoel
What induces thickening of ectoderm into a neutral plate
chordamesoderm
ectomesenchyme, considered as the 4th germ layer which arise from ectoderm forms cartilage and bones of the head, pharyngeal cartilages; peripheral nerve ganglia, some glandular tissues; melanocytes
neural crest cells
ectodermal cells
neural plate
formed from neural plate; folds of cells that arches and meet at the mid-dorsal line and forms the neural tube
neural folds
– encloses the neurocoel (cavity)
neural tube
anlagen of the dorsal hollow nerve cord
neurocoel
migration of primordial germ cells
organogenesis
establishes the head and tail
holoblastic embryo
– 3 germ layers spread faced down on the uncleaved yolk
Type of embryo
meroblastic embryo
embryo increase in ____ while anchenteron becomes part of the ____ tube
length
gut
enterocoelom (blastopore) becomes anus while the mouth opens anteriorly
schizocoely
____form the lining of the gut
endoderm
major structure formed by dermatome (outer epimere)
skin dermis
major structure formed by middle epimere
muscles
major structure formed by inner epimere
vertebral column
major structure formed by chordamesoderm
notochord
major structure formed by intermediate mesoderm
kidney, urogenital ducts
major structure formed by somatic hypomere
bones
major structure formed by splanchnic hypomere
blood, heart, gut, smooth muscle, visceral perotineum
major structure formed by somatic endoderm
skin epidermis, teeth enamel, stomodeum, proctodeum
major structure formed by neural plate ectoderm
brain, spinal cord
major structure formed by neural plate ectoderm
brain, spinal cord
major structure formed by epidermal placodes
capsules
major structure formed by ectomesenchyme
spinal ganglia, nenurocranium, aortic arches, heart septum
in them, yolk cleaves and directly incorpotaed into somatic cells
amphibians
➢
Yolk supply is limited
➢
No fetal membranes required to sustain larva
➢
Early hatching > larva
➢
Example include caecilian, frogs, newts, salamanders
amphibians
in them, yolk remains uncleaved, eggs are laid in water; yolk sac is the only fetal membrane found
fishes
➢
Yolk sac – formed from 3 germ layers to absorb the yolk into the body
➢
Respiration and excretion are direct contract with environment
fishes
in them, there is a primitive streak, extraembryonic, mesoderm splits, forms extraembryonic coelom, splanchnoleure
reptiles and birds
Splanchnopleure + adjacent yolk ++
yolk sac
– under the shell; near the chorioallantoic membrane
allantois
Respiration, receives excretory wates; absorbs albumen takes some minerals from the shell
allantois
Somatopleure + head fold of amnion =
chorion
amnion
“water bag” contains amniotic fluid that bathes the embryo; cushions the embryo
amnion
nourish their young in uterus, exchange and nutrition of fetus occurs between fetal and maternal bloodstream by placenta
placental mammals
vascularized (blood vessels) supplies nutrition to placenta
yolk sac and allantois
avascular (no blood vessels) cannot support placenta
chorion and amnion
homologous to umbilical circulation (mammals)
allantoic circulation
sac filled with amniotic fluid bathes the fetus; allows the early fetus to move freely and protects the fetus from pressure of maternal abdomen
amnion
collective term for reptiles, birds and mammals because the possess amnion
amniotes
chorion (lies between allantois and uterus) incorporated into the placenta, chorioallantoic membrane
maternal contribution
allantois (in most mammals) as fetus grows, allantois decreases in size
fetal contribution
egg laying mammals, deposit eggs in pouch
protherians
marsupials (pouched mammals) no typical placenta
metatherians
includes the skin and the related structures that cover and protect the bodies of animals
integumentary system
the integument of these group of animals includes shells and exoskeletons as body covering
invertebrates
the integument of these group of animals include skin, scales, feathers, hair, and glands
vertebrates
its integument is made up of skin that includes glands, hair, and nails
human
in humans, this protects the body, prevents, water loss, regulates body temperature, and senses the external environment
skin
Evolutionary adaptations of the integumentary system:
regulation of body temp
excretion of waste materials
vitamin D formation via UV radiation
reception of environmental stimuli
locomotion
movement of nutrients and gas
outer covering of the body of vertebrates
integument
Commonly referred to as skin
integument
Include the mucous membrane lining of the mouth, eyelids, nostrils, and the openings of rectum and urogenital organs
integument
Forms the interface between organisms and the external environment
integument
Integument consists of two layers
epidermis
dermis
between the epidermis and dermis this lies
basal membrane
underlying support where epithelial cells are rested
basement membrane
Demarcates the underlying connective tissue from epithelium
basement membrae
membrane proteins of the epithelial cells are anchored in basal lamina, has two 2 major glycoproteins
basal lamina
▪
Laminin
▪
Type IV collagen
▪
Acts as selectively permeable filter between epidermis and dermis connective tissue
basal lamina
reticular fibers embedded in ground substance that connect the basal lamina with the underlying CT
reticular lamina
*r – layer under the dermis that is made up of very loose connective and adipose tissue
Hypodermis or subcutaneous laye
what is converted to vitamin D in skin
cholesterol
– arises from ectoderm
epidermis
attached to basement membrane, it actively cell divides to replenish the outer periderm
stratum germinativum
Differentiates into stratified layer with mucous or keratin coat on the outer surface (prevent desiccation)
epidermis
origin varies (layer of skin)
dermis
principal origin, from the outer wall of dermomyotome of somites
dermatome
form dermatome that settles under the epidermis, some may have stratum compactum that have diffuse, irregular collagen bundles
connective tissue of dermis
beneath the dermis, forms the fascia composed of loose connective tissue and adipose tissue
hypodermis
provides waterproofing and serves as a barrier to infection
epidermis
serves as a location for appendages of skin
dermis
called the basement membrane
hypodermis
composed of epithelial cells derived from ectoderm
epidermis
Serve as the interface between the organism and environment
epidermis
Made up of stratified squamous layer of epithelial cells
epidermis
in them, epidermis is covered by a thin coat of mucus and contains unicellular glands
aquatic vertebrates
in them, epidermis is covered by a dead, water resistant cornified cells (s. corneum)
terrestrial vertebrates
outermost; composed of dead cells; keratinized
stratum corneum
Keratinized layer of skin responsible for keeping water in the body and keeping other harmful chemical and pathogens out, making skin a natural barrier to infection
stratum corneum
translucent layer, composed of flat, dead and non-nucleated cells
stratum lucidum
– composed of keratohyaline granules
stratum granulosum
prickle cell layer; composed of Langerhans cells
stratum spinosum
inner layer, composed of actively dividing cells
stratum germinativum
90 percent of epidermal cells; produce keratin
keratinocytes
8 percent of epidermal cells; produce melanin
melanocytes
capable of trapping antigen in the skin
langerhans cells
make contact with the ending of a sensory neuron
merkel cells
lies below the epidermis and contains an umber of structures including blood vessels, nerves, hair follicles, smooth muscle, glands, and lymphatic tissue
dermis
Consist of loose connective tissue otherwise called areolar connective tissue – collagen, elastin, and reticular fibers are present
dermis
Provides tensile strength and physiologic support for the interfacing epidermis
dermis
Has an ancient and persistent potential to form bone
dermis
also known as hypodermis, not part of the skin, lies below the dermis
subcutaneous layer
Purpose is to attach the skin to the underlying bone and muscle as well as supplying it with blood vessels and nerves
subcutaneous layer
Consist of loose connective tissue and elastin
hypodermis