Lecture Exam #4 Ch. 34 Flashcards
where do the animals called vertebrates get their name from?
vertebrae, the series of bones that make up the backbone
what modern animals did vertebrates give rise to? (3) (ARM)
1) amphibians
2) reptiles (including birds)
3) mammals
how many vertebrates are there on Earth and what do they include?
57,000 and they include some of the largest organisms
what do verterbrates have?
great disparity with a wide range of differences within the group
what do chordates have?
a notochord and a dorsal, hollow nerve cord
what type of animals are chordates and what clade do they belong to?
they are bilaterian and they belong to the clade konwn as dueterostomia
what does chordates comprise?
1) all vertebrates
2) group of invertebrates, the urochordates and cephalochordates
what do all chordates share?
a set of derived characteristics
what do some species of chordates have?
some of the derived traits only during embryonic development
4 key characters of chordates (NDPM)
1) notochord
2) dorsal, hollow nerve cord
3) pharyngeal slits or clefts
4) muscular, post-anal tail
a longitudinal, flexible rod between the digestive tube and nerve cord
notochord
what does the notochord provide?
skeletal support throughout most of the length of the chordate
in most verterbrates what devleops
a more complex, jointed skeletal and the adult retains only the remnants of the embroynoic notochord
what does the nerve cord of a chordate embryo develop from?
a plate of ectoderm that rolls into a tube dorsal to the notochord
what does the nerve cord develop into?
the central nervous system (the brain and the spinal cord)
found in most chordates, grooves in the pharynx that develop into slits that open to the outside of the body
pharyngeal clefts
functions of the pharyngeal slits (3) (SGD)
1) suspension- feeding structures in many inveterbrate chordates
2) gas exchange in vertebrates (except vertebrates with limbs, the tetrapods)
3) develop into parts of the ear, head and neck in tetrapods
in many chordate species what is greatly reduced during embryonic development?
the tail
what does the tail provide in many aquatic animals?
propelling force
what are lancelets (cephalochordata) named for?
their bladelike shape
what type of feeders are lancelets?
marine suspension feeders
what do lancelets retain as adults?
the characteristics of the chordate body plan
what is more closely related to chordates than lancelets?
tunicates
when does tunicates most resemble chordates?
during their larval stage (which may only last a few min)
what does a tunicate do as an adult?
draws in water through an incurrent siphon, filtering food particles
what do tunicates or “sea squirts” do when attacked?
shoot water through their excurrent siphon
what may have ancestral chordates resembled?
lancelets
where are the same hox genes that organize the vertebrate brain expressed in as well
the lancelets’s simple nerve cord tip
what does the sequencing of the tunicate genome indicate? (2) (GG)
1) genes associated with the heart and thyroid are common to all chordates
2) genes associated with transmission of nerve impulses are unique to vertebrates
what two essential tasks does the skeletal system and nervous system allow vertebraes to do? (2) (CE)
1) capture food
2) evade predators
how many sets of hox genes does vertebrates have?
two or more
how many hox genes do lancelets and tunicates have?
only one cluster
derived characteristics of vertebrates (VEF) (3)
1) vertebrae enclosing a spinal cord
2) an elaborate skull
3) fin rays, in aquatic forms
what does fossil evidence show that the earliest vertebrates lacked?
jaws
what two lineage of jawless vertebrates remain?
hagfishes and lampreys
what do members of hagfishes and lampreys lack?
a backbone
what indicates that hagfishes and lampreys are vertebrates?
the presence of rudimentary vertebrae and the results of phylogentic analysis
what clade of living jawless vertebrates do the hagfishes and and lampreys form together?
cyclostomes
what much larger clade do vertebrates with jaws make up?
gnathostomes
characterisitcs of hagfishes (4) (JCRF)
1) jawless vertebrates
2) have a carliganious skull
3) reduced vertebrae
4) flexible rod of cartlage derived from the notochord
what small features do hagfishes have?
small brain, eyes, ears and tooth-like formations
what type of animals are hagfishes?
marine, most are bottom-dwelling scavengers
how do the parasites, lampreys (Petromyzontida) eat?
ones that feed by clamping their mouth onto live fish
what type of environments do lampreys inhabit?
various marine and freshwater habitats
what does lampreys have?
cartilaginous segments surrounding the notochord and acrhing partly over the nerve cord
what documents the transitions to craniates?
fossils from the cambrian explosion
what were the most primitive of the fossils from the Cambrian evolution?
those of the 3-cm-long Haikouella
characteristics of the Haikouella (4) (WEMN)
1) well-formed brain,
2) eyes
3) muscular segments
4) no skull or ear organs
what were among the earliest verterbrates in the fossil record, dating from 500 to 200 million years ago?
conodonts
what did condodonts have?
mineralized skeletal elements in their mouth and pharynx
what was common the fossil record of condodonts?
fossilized dental elements
what are other groups of jawless vertebrates armored with?
defensive plates of bone on their skin
what did minerlization of bone and teeth appear to have originated with?
vertebrate mouthparts
what became fully mineralized much later?
the vertebrate endoskeleton
today what outnumbers jawless vertebrates?
jawed vertebrates, gnathostomes
what do gnathostomes include? (6) (SRLARM)
1) sharks and their relatives
2) ray-finned fishes
3) lobe-finned fisehs
4) amphibians
5) reptiles (including birds)
6) mammals
what are gnathostomes named for?
their jaws
what does the jaws with the help of teeth in gnasthostomes help do?
grip food items firmly and slice them
what are the jaws of gnasthostomes hypothesized to have evolved by?
modificiation of skeletal rods that supported pharyngeal (gill) slits s
other characteristics common to gnathostomes (3) (GEA)
1) genome duplication including duplication of hox genes
2) an enlarged forebrain associated with enhanced smell and vision
3) in aquaitc gnathostomes the lateral line system, which is sensitive to vibration
what are the earliest gnasthostomes in the fossil record that appeared 440 million years ago?
an extinct lineage of armored verterbrates called placoderms
what is another group of jawed vertebrates called that radiated during the Silurian and Devonian periods (444 to 359 million years ago)?
acanthodians
what are three lineages of jawed vertebrates that survive today? (3) (CRL)
1) chondrichthyans
2) rayed-finned fisehs
3) lobe-fins
what do chondrichthyans include?
sharks, rays and their relatives
what do chondrichthyans have?
a skeleton composed primarily of cartlage
what does the largest and most diverse group of chdonrichtyans include?
sharks, rays and skates
what is a second subclass of chondorichythans composed of?
a few dozen species of ratfishes or chimaeras
what type of bodies do sharks have and what type of swimmers are they?
a streamlined body and are swift swimmers
what type of feeders are the largest sharks and what are most of them?
suspension feeders but most are carnivores
what type of digestive system do sharks have?
a short digestive tract with a ridge called the sprial valve
what does the spiral valve do for sharks’ digestive system?
increases the digestive surface area
what type of acute senses do sharks have?
sights, smell and the ability to detect electrical fields from nearby animals
how are sharks eggs fertilized?
internally but embryos can develop in different ways
different ways that embryos can develop in sharks (3) (OOV)
1) oviparous
2) ovoviviparous
3) vivparous
eggs hatch outside the mother’s body
oviparous
the embryo develops within the uterus and is nourised by the egg yolk
ovoviparous
the embryo develops within the uterus and is nourished through a yolk sac placenta from the mother’s blood
vivparous
in sharks, what does the reproductive tracts, excretory system and digestive tract empty into?
a common cloaca
today, what are sharks severly threatened by?
overfishing (pacific population plummeted upto 95%)
what clade of gnasthostomes does the vast majority of vertebrates belong to?
osteichthyes
what do nearly all living osteichthyes have?
a bony endoskeleton
what does osteichthyans include?
bony fishes and tetrapods
what are aqautic osteichthyans?
vertebrates we informally call fishes
how do most fishes breath?
by drawing water over gills protected by an operculum
how do fishes control their buoyancy?
with an air sac known as a swim bladder
what do fishes have?
a lateral line system
what type of embryo development do most fishes have?
oviparous but some have internal fertilization and birthing
what does actinopterygii the ray-finned fishes include?
nearly all the familiar aquaitc osteichthyans
when did ray-finned fishes originate?
during the Silurian period
what are the fins of ray-finned fishes supported mainly by long, flexible rays modified for?
maneuvering, defense and other functions
what did industrial-scale fishing operations drive many ray-finned fish populations to do?
collapse
what are populations of ray-finned fishes also affected by and what does it change?
dams, and it changes water flow patters, affecting prey capture, migration and spawning
when did lobe-fins (sacropterygii) originate?
during the Silurian period
what do lobe-fins have and for what?
muscular pelvic and pectoral fins to swim and “walk” underwater across the substrate
what are the 3 lineages of lobe-fins that survive? (3) (CLT)
1) coelacanths
2) lungfishes
3) tetrapods
even though Coelacanths were thought to have become extinct 75 million years ago, what was caught?
a living coelacanth caught of the coast of South Africa in 1938
where are the living lungishes all found?
in the Southern Hemisphere
though gills are main organs for gas exhange in lungfishes what else can they do?
surface to gulp air into their lungs