digestive anatomy of RUMINANT Flashcards
upper incisors
none, replaced by a hard, fibrous dental pad
what separates the incisors from cheek teeth
diastema
cheek teeth (pre molars and molars) are
incisors are
hypsodont
brachydont
Cheek teeth in herbivores have a complex pattern on the surface due to
invaginations of the enamel layer
As the tooth surface wears down over time, ____ occur on the surface
hard enamel crests; help grind up food
Upper incisors replaced by ____ in ruminants
dental pad
typical foot and mouth disease is seen in
ulceration of the dental pad
dental formula for young and for adults
young:
2(Di 0/4 Dc 0/0 Dp 3/3) = 20
adult:
2(I0/4 C0/0 P3/3 M3/3) = 32
*canine and 4th incisor interchangeable
eruption date for cattle and sheep/ goats
- cattle:
- incisor 1; 1.5-2yrs
- incisor 2; 2-2.5 yrs
- incisors 3; 3-3.5 yrs
- incisors 4; 3.5-4 yrs
- sheep
- incisor 1; 1-1.5 yrs
- incisor 2; 1.5-2 yrs
- incisor 3; 2.5-3 yrs
- incisor 4; 3-4 yrs
initial mastication vs re-mastication or rumination
- initial; quick
- remastication or rumination; slower, plants ground between cheek teeth
lateral chewing movement is due to
- strong masseter and
pterygoid muscles, - relatively narrow, though long mandibles,
- the slope, and roughened surface of the cheek teeth.
tongue is highly
extensible to grasp food
lower incisors cut off grass against
dental pad
Small ruminants have highly mobile ___ which enables them to nibble and crop plants close to the ground
lips
small ruminants have a cleft upper lip known as the
philtrum
negative pressure piston-like action of tongue is aided by
torus; prominence over the caudal part of the body and root
torus has a _______ on its rostral side
transverse lingual fossa
conical papilla
inner surface of lips and cheeks, root of tongue large, pointing backward backward filiform papillae , mechanical in function
lenticular papillae
flat, rounded, mechanical, mainly on torus, act like suction cups
vallate papilla aka circumvallate papillae
very large, on dorsal root of tongue, V shape array, surrounded by deep groove with taste buds on the side
foliate papilla
may be absent, lateral root of tongue, taste buds
true or false the mandible can movie in all 3 planes
true
lateral movement is due to what muscles
strong masseter and pterygoid muscles
when digastric muscle contracts
jaw opens
cattle produce up to ___L saliva a day
150
what is main source of liquid in rumen
saliva
saliva functions
- form food bolus
- provide alkaline buffer to rumen and reticulum
- maintain pH in rumen (contains large quantities of buffer)
- essential for fermentation
large ____ gland covers the more medial ____ gland
parotid
mandibular
the duct of parotid gland runs ventral to the
masseter muscle
the dorsal buccal branch of facial nerve passes over
masseter muscle
the ___ duct passes over the facial artery and nerve
parotid
The
ventral end of the ______ forms a conspicuous swelling in the
intermandibular space which may be mistaken for the mandibular lymph node, although the salivary gland is larger, softer and more rostral.
mandibular salivary gland
what contracts during rumination
what relaxes
rumen and reticulum
cardiac sphincter relaxes
during rumination when the rumen and reticulum contract, the fluid level is raised to the
entrance of the esophagus
how do rumens get the food up during rumination
- rumen and reticulum contract –> fluid raised to entrance of esophagus
- cardiac sphincter relaxes
- cow inspires on closed glottis–> lowers thoracic pressure, esophagus acts like vacuum
- cud passes into it
- cud passes to mouth by reverse peristalsis
- swallows excess fluid
- cud chewed (remasticated) mixed w more saliva
- swallowed again
nasopharynx divided dorsally by the
median pharyngeal septum
the ox has palatine tonsils which ahve follicles that open into _____. These then open into a deep common _____
crypts
sinus tonsillaris
, in most species, the oesophagus decreases in diameter as it approaches the stomach, and the thickness of the muscular wall increases. In ruminants:
the reverse occurs: the overall size of the oesophagus increases, and the wall becomes thinner as the oesophagus approaches the cardia.
muscle type in tunica muscularis
100% skeletal
the mucosa of the esophagus is particularly ____ in the ruminant
cornified
in ruminants fermentation takes place before or after reaching the glandular stomach and the small intestines
before
Fermenting herbivores share a mutually beneficial symbiotic relationship with ______ microorganisms
anaerobic
what do the microorganisms provide that can be used as energy
metabolites- VFAs
assimilation
the process by which living organisms convert food into vitamins, minerals, and other chemicals that the body needs in order to function
In foregut fermenters, this fermentation process takes place ____ the food reaches the glandular stomach, and the small intestines
before
ruminant digestion has led to enlargement of what parts of stomach
cranial non-glandular stomach
gases produced from rumination are removed by
eructation (burping)
rumen stomach
- rumen
- reticulum
- omasum
- abomasum (true stomach)
which stomachs make up the non-glandular stomach where foregut fermentation takes place
rumen, reticulum and omasum
what is the glandular stomach of the ruminants
abomasum
abomasum: rumen/reticulum ration at birth, 2 months and adult
birth 2:1
2 months 1:1
adult 1:9
rumen develops as an outgrowth of the
fundus of stomach, gorws caudal and dorsal (back and over the rest of the stomach which then becomes abomasum)
reticulum develops as an outgrowth of
rumen
omasum develops drom the
lesser curvature of primitive stomach
ruminant stomach development from primitive stomach summary
- rumen outgrowth from fundus
- reticulum outgrowth of rumen
- omasum growth from lesser curvature
- abomasum is rest of stomach
rumen lies on
LHS of abdominal cavity
location of parts of stomach
- reticulum lies cranially against the diaphragm
- omasum lies on RHS cranially
- abomasum lies ventrally
- rumen on LHS
rumen parietal surface is ____
visceral surface is ____
right
left
part of dorsal surface of rumen is attached to
body wall
rumen is divided into __ sacs
6
the 6 sacs of rumen are divided by deep grooves containing
fat, blood vessels, lymphatics, and lymph nodes and nerves
covered by visceral peritoneum
shallow left and right longitudinal grooves are connected cranially and caudally by the
deep cranial and caudal grooves
sacs of rumen: caudal end
The dorsal blind sac and the ventral blind sac are marked off by the dorsal
and the ventral coronary grooves. The blind sacs are separated by the caudal groove which runs left to right. Again, we’ve got a couple of coronary grooves, that go both dorsal and ventral. So they’re named the dorsal coronary groove, and the ventral coronary groove. These demarcate the dorsal and ventral blind sacs
sacs of rumen; cranial end
The dorsal and ventral sacs have bulges at the cranial end; these form the cranial sac, and the ventral recessus ruminis. Again, off
the cranial end of the dorsal and ventral sacs, there’s a cranial sac, and ventrally
there’s a sac called the recessus ruminis. The cranial pillar separates the cranial sac and the recessus ruminis.
cranially, the cranial sac of rumen communicates with the reticulum via the wide
ruminoreticular opening: opening from rumen into reticulum
also where the esophagus enters at the cardia
opposite the cardia is the deep
ruminoreticular groove
ruminal grooves form internal
pillars
separating the rumen and reticulum is the high U shaped
ruminoreticular fold
lining of the rumen internally is what kind of epithelium
keratinized, stratifed squamous epithelium which is thrown up into papillae
the papillae on the inside lining of rumen function
increase SA for transfer of water, small solutes and heat
which areas of internal rumen are the papillae most prominent
where are they absent/ reduced
cranial, ventral and blind sacs
reduced/absent in over the pillars and on part of the dorsal sac
mucosa of the complex stomach is often
folded; increase SA
rumen lamina propria
irregular dense CT, containing capillaries and nerves
lymphoid tissue ABSENT
muscularis mucosa of the rumen
ABSENT
submucosa of the rumen
loose CT
tunica muscularis of the rumen
how many layers, where is it thick
2-3 layers, thick at the pillars
inner circular layer, outer longitudinal layer and also a third layer of internal oblique muscles; only found in rumen and reticulum
where does the tunica muscularis have a third layer of internal oblique muscles, and why
rumen and reticulum,; helps contract and raise fluid level to esophagus for rumination
serosa of rumen has what epithelium
simple squamous or cuboidal or mesothelial cells where is is contunous with the visceral periotneum
part of rumen that attaches to the abdominal roof has
tunica adventitia instead of tunica serosa
the subserosa of the rumen is rich in
adipose tissue
mesothelial cells secrete small amounts of
serous fluid; peritoneal fluid (allows organs to slip past eachother)
the mesothelium layer of the tunica serosa of rumen is continuous with the
peritoneum; borders peritoneal cavity
bloat
ruminal distension from gases of fermentation