Hearing: The Auditory System, Localisation, And Auditory Scene Analysis Flashcards
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Acoustic shadow
an area through which sound waves fail to propagate, due to topographical obstructions or disruption of the waves via phenomena such as wind currents, buildings, or sound barriers
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Additive synthesis
process of adding harmonics to create complex sounds
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Amplitude
the fluctuation or displacement of a wave from its mean value
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Amplitude modulation
the modulation of a wave by varying its amplitude, used especially as a means of broadcasting an audio signal by combining it with a radio carrier wave
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Amplitude-modulated noise
the detection of modulation versus the detection of modulation rate
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Aperiodic sounds
sound whose pattern does not repeat itself as do those of the periodic sounds
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Attack
changes occurring before the sound reaches its steady-state intensity
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Audibility curve
the relationship between the threshold of hearing for a pure tone, expressed in decibels sound-pressure level (dB SPL), and the frequency of the tone
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Auditory canal
canal leading from the opening of the external ear to the eardrum
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Auditory localisation
describes an organism’s ability to perceive the location of the source of a sound
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Auditory
falls between the audibility curve and and the. threshold for feeling
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Auditory scene analysis
a proposed model for the basis of auditory perception. This is understood as the process by which the human auditory system organizes sound into perceptually meaningful elements
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Auditory space
surrounds an observer and exists wherever there is sound
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Auditory stream segragation
the perceptual grouping of sounds, to form coherent representations of objects in the acoustic scene, and is a fundamental aspect of hearing and speech perception
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Azimuth
the direction of a celestial object from the observer, expressed as the angular distance from the north or south point of the horizon to the point at which a vertical circle passing through the object intersects the horizon.
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Basilar membrane
a stiff structural element within the cochlea of the inner ear which separates two liquid-filled tubes that run along the coil of the cochlea, the scala media and the scala tympani.
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Bass ratio
ratio of low to middle frequencies reflected from surfaces
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Belt area
the secondary auditory cortex immediately surrounding the core
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Binaural cues
any difference in the sound arriving at the two ears from a given sound source (interaural difference) that acts as a cue to permit auditory localization
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Characteristic frequency
The frequency of a sound at which the threshold of a single fibre of an auditory nerve is lowest and to which it is therefore most responsive
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Cilia
the mechanosensing organelles of hair cells, which respond to fluid motion in numerous types of animals for various functions, including hearing and balance
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Cochlea
a spiralled, hollow, conical chamber of bone, in which waves propagate from the base (near the middle ear and the oval window) to the apex (the top or center of the spiral)
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Cochlear amplifier
a positive feedback loop within the cochlea that amplifies the traveling wave
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Cochlear nucleus
the obligatory nucleus in which all ascending information from the cochlea forms synaptic connections with the auditory brain
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Coincidence detectors
the context of neurobiology is a process by which a neuron or a neural circuit can encode information by detecting the occurrence of temporally close but spatially distributed input signals
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Condensation
part of a sound wave where the density is higher than normal
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Decay
rate at which the music fades to silence
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Direct sound
sound that reaches the listener’s ears straight from the source
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Effect of the missing fundamental
Removal of the first harmonic in a sound with the same perceived pitch, but with a different timbre
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Event related potential
the measured brain response that is the direct result of a specific sensory, cognitive, or motor event
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First harmonic
another term for the fundamental frequency which sets the repetition rate
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Frequency
the number of occurrences of a repeating event per unit of time
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Frequency spectrum
display of harmonics of a complex sound
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Fundamental
the lowest frequency of a periodic waveform
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Fundamental frequency
the lowest frequency which is produced by the oscillation of the whole of an object, as distinct from the harmonics of higher frequency
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Hair cells
the sensory receptors of both the auditory system and the vestibular system in the ears of all vertebrates, and in the lateral line organ of fishes
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Harmonic
a signal or wave whose frequency is an integral (whole-number) multiple of the frequency of some reference signal or wave
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Hertz (Hz)
the derived unit of frequency in the International System of Units (SI) and is defined as one cycle per second
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Incus
a bone in the middle ear. The anvil-shaped small bone is one of three ossicles in the middle ear
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Indirect sound
sound that is reflected off of environmental surfaces and then to the listener
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Inferior colliculus
the principal midbrain nucleus of the auditory pathway and receives input from several peripheral brainstem nuclei in the auditory pathway, as well as inputs from the auditory cortex
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Inner hair cells
the sensory cells of the cochlea, are responsible for signal transduction
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Interaural level difference (ILD)
the difference in loudness and frequency distribution between the two ears
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Interaural time difference (ITD)
the difference in arrival time of a sound between two ears
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Intimacy time
time between when sound leaves its source and when the first reflection arrives
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Jeffress model
a neurocomputational model that explains how auditory systems can register and analyze small differences in the arrival time of sounds at the two ears in order to estimate the direction of sound sources in the azimuthal plane.
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Loudness
the subjective perception of sound pressure
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Malleus
a hammer-shaped small bone or ossicle of the middle ear which connects with the incus and is attached to the inner surface of the eardrum
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Medial geniculate nucleus
part of the auditory thalamus and represents the thalamic relay between the inferior colliculus (IC) and the auditory cortex (AC).
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Melody
a sequence of single notes that is musically satisfying; a tune.
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Middle ear
the portion of the ear internal to the eardrum, and external to the oval window of the inner ear
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Middle ear muscles
two muscles which serve a protective function in the middle ear; the tensor tympani and stapedius
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Monaural cue
relies on sounds reaching a single ear to constrain the set of possible sound sources
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Octave
the interval between one musical pitch and another with double its frequency
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Onset time
sounds that start at different times are likely to come from different sources
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Organ of Corti
the receptor organ for hearing and is located in the mammalian cochlea. This highly varied strip of epithelial cells allows for transduction of auditory signals into nerve impulses’ action potential.
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Ossicles
three bones in either middle ear that are among the smallest bones in the human body.
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Outer ear
the external portion of the ear, which consists of the auricle (also pinna) and the ear canal
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Oval window
a membrane-covered opening that leads from the middle ear to the vestibule of the inner ear
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Periodic sounds
sounds which repeats itself at regular intervals
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Periodicity pitch
Removal of the first harmonic in a sound with the same perceived pitch, but with a different timbre
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Phase locking
the mechanism by which the peripheral auditory system keeps track of the times of occurrence of the ongoing amplitude fluctuations in sounds
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Pinnae
the external part of the ear in humans and other mammals; the auricle
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Place theory
a theory of hearing that states that our perception of sound depends on where each component frequency produces vibrations along the basilar membrane
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Precedence effect
a binaural psychoacoustical effect. When a sound is followed by another sound separated by a sufficiently short time delay (below the listener’s echo threshold), listeners perceive a single auditory event
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Presbycusis
the loss of hearing that gradually occurs in most individuals as they grow older
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Primary auditory cortex (A1)
part of the temporal lobe that processes auditory information in humans and many other vertebrates
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Pure tone
a sound with a sinusoidal waveform; that is, a sine wave of any frequency, phase, and amplitude
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Rarefaction
reduction in the density of something, especially air or a gas
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Resolved harmonics
lower harmonics that are separated out in the cochlea (i.e., they excite distinct places on the basilar membrane)
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Resonance
the phenomena of amplification that occurs when the frequency of a periodically applied force is in harmonic proportion to a natural frequency of the system on which it acts
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Resonant frequency
a natural frequency of vibration determined by the physical parameters of the vibrating object
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Reverberation time
the time it takes sound to decrease by 1/1000th of its original pressure
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Scale illusion
an auditory illusion in which principles of grouping by frequency, proximity, and spatial location are put into conflict and in which frequency proximity wins out
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Sound
vibrations that travel through the air or another medium and can be heard when they reach a person’s or animal’s ear.
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Sound pressure level (SPL)
the pressure level of a sound, measured in decibels (dB)
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Sound wave
a wave of compression and rarefaction, by which sound is propagated in an elastic medium such as air.
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Spaciousness factor
fraction of all the sound received by listener that is indirect
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Stapes
a small stirrup-shaped bone in the middle ear, transmitting vibrations from the incus to the inner ear.
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Subcortical structures
a group of diverse neural formations deep within the brain which include the diencephalon, pituitary gland, limbic structures and the basal ganglia.
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Superior olivary nucleus
a group of nuclei located in the brainstem near the junction of the pons and medulla. It is the first auditory relay after the cochlear nucleus on the way to the auditory cortex and is the major point at which information from the two ears is integrated.
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Tectorial membrane
an extracellular connective tissue that covers the mechanically-sensitive hair bundles of the sensory receptor cells in the inner ear.
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Temporal coding
a type of neural coding which relies on precise timing of action potentials or inter-spike intervals
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Timbre
the character or quality of a musical sound or voice as distinct from its pitch and intensity.
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Tip links
extracellular filaments that connect stereocilia to each other or to the kinocilium in the hair cells of the inner ear
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Tonality
the arrangement of pitches and/or chords of a musical work in a hierarchy of perceived relations, stabilities, attractions and directionality
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Tone height
perceptual experience of increasing pitch as tone’s frequency (wavelength increases)
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Tympanic membrane
a membrane forming part of the organ of hearing, which vibrates in response to sound waves. In humans and other higher vertebrates it forms the eardrum, between the outer and middle ear.