Chapter 16 Lec. 1 Flashcards
The earliest life
Earth is 4.5 billion years old.
Evidence for microbial life can be found in rocks 3.86 billion years old.
Evidence
Biomarkers. Isotopes -Carbon 12 6p and 6n -Carbon 13 6p and 7n -Carbon 14 6p and 8n
Stromatolites
Fossilized microbial mats consisting of layers of filamentous prokaryotes and trapped sediment.
Found in rocks 3.5 billion years old.
Comparisons of ancient and modern stromatolites
-Anoxygenic
-Oxygenic
Origin of cellular life
Early earth was anoxic and hot.
First biochemical compounds were made by abiotic systems that set the stage.
-Amino acids
-Nucleic acids
MIller-Urey Experiment (1953)
Academic “grand father”.
Created early earth atmosphere.
-Formed amino acids, nucleic acids, etc…
surface origin hypothesis
First membrane enclosed, self replicating cells arose out of primordial soup in ponds on earth’s surface.
Dramatic temperature fluctuations and mixing from meteor impacts, dust clouds and storms argue AGAINST this hypothesis.
Subsurface origin hypothesis
Life originated at hydrothermal springs on ocean floor.
- Conditions more stable.
- Steady and abundant supply of energy (H2 and H2S).
RNA world theory
Prebiotic chemistry of early earth set stage for self replicating systems.
First self replicating systems may have been RNA based.
RNA can bind small molecules.
-RNA has catalytic activity; may have catalyzed its own synthesis.
-RNA is an informational molecule.
Transition to DNA
DNA is a more stable molecule and became genetic repository.
Proteins took over catalysis jobs.
Three part systems evolved and became universal.
-DNA
-RNA
-Protein
Lipid Bilayer
Buildup of lipids.
Can naturally form bilayer.
Enclosed RNA became self replicating.
Last universal common ancestor (LUCA)
Populations of early cells from which cellular life may have diverged into ancestors of modern bacteria and archaea.
Characteristics of LUCA
Early earth was anoxic, thus energy generating metabolism of primitive cells: -anaerobic -chemolithotrophic -Obtained carbon from CO2 Likely: -Obtained energy from H2 -Generated by FeS reacting with H2S or UV light. -Thermophilic
Chemoorganotrophy
Early life forms were chemolithotrophic.
-Produce large amounts of organic compounds.
Organic material provided abundant, diverse and continually renewed source of reduced organic carbon.
-stimulating evolution of chemoorganotrophic metabolisms.
Archaea and Bacteria
Evidence suggests ancestors of bacteria and archaea diverged 4 billion years ago.
As lineages diverged, distinct metabolisms developed.
Development of oxygenic photosynthesis changed course of evolution.
Oxygenic Photosynthesis
2.7 billion years ago cyanobacteria invented oxygenic photosynthesis.
Rise of O2
- 4 billion years ago.
- Ocean absorbed O2 for ~300 million years.
Microbial diversification
Development of oxic atmosphere.
-Evolution of new metabolic pathways
Oxygen spurred evolution of organelle containing eukaryotic microorganisms.
-eukaryotic fossils are around 2 billion years old.
-multicellular and more complex eukaryotes found 1.9-1.4 billion years ago.
The global carbon cycle
Volcanism and weather.
-Volcanism: CO2 gas in atmosphere.
Weathering
Breakdown of minerals.
-Takes carbon into core.
Volcanism vs. weathering
Temperature is controlled by weathering.
-Carbon in the atmosphere.
Endosymbiosis
Supported hypothesis for origin of eukaryotic cells.
Contends that mitochondria and chloroplasts arose from symbiotic association of prokaryotes within another type of cell.
Endosymbiotic origin of eukaryotes
- Eukaryotes began as nucleus bearing lineage that later acquired mitochondria and chloroplasts by endosymbiosis.
- Eukaryotic cells arose from intracellular association between O2 consuming bacteria which gave rise to mitochondria and an archaea.
Endosymbiotic origin 2
Eukaryotic cells are chimeric.
Supported by:
-eukaryotes have similar lipid and energy metabolisms to bacteria.
-transcription and translational machinery similar to archaea
The evolutionary process
Mutations
-Changes in nucleotide sequence of an organisms genome.
-Occurs because of errors in replication, UV radiation.
-Adaptive mutations improve fitness of an organism, increasing its survival
Other genetic changes include gene duplication, horizontal gene transfer and gene loss.