Lecture 12 Flashcards
phylum anthophyta
angiosperms all in one phylum, vast majority of seed plants
flower
a specialized shoot with up to 4 whorls of modified leaves
4 parts of flower
- sepals (sterile)
- petals (sterile)
- stamens (anther + filament) - microsporophylls
- carpels (stigma + stile + ovary) - megasporophylls
ovary
base of carpel, contains 1 or more ovules
perfect flower
contains both stamens and carpels
imperfect flower
contains only stamens or carpels
monoecious
imperfect male and female flowers on same plant
dioecious
imperfect male and female flowers on different plants
animal pollination
angiosperms are more reliant on animal pollinators, flowers modified to attract and dispense pollen, flowers provide food rewards (sugary fluids ie. nectar, or pollen itself)
fruit
mature ovary containing seeds - the ‘container’ part of angiosperm, may be dry and thin or thick and fleshy, aid dispersal of seed
cotyledons
embryonic seed leaves, present inside seed along with embryonic root (radicle), can have 1 or 2
monocots and dicots
previous subdivision of Anthophyta based on 1 or 2 cotyledons
angiosperm life cycle
- anthers produce microspores that produce male gametophytes (pollen grains)
- ovules produce megaspores that produce female megagametophytes (embryo sac)
- pollen lands on stigma (pollination)
- pollen germinates
- double fertilization
pollen grain 2 cells
- generative cell –> sperm
2. tube cell –> pollen tube
embryo sac
megagametophyte in angiosperms, contains only 7 cells (less than gymnosperms) and 8 nuclei
polar nuclei
one cell in embryo sac contains 2 nuclei
pollen germinates
generative cell divides to form 2 sperm cells, pollen tube grows down style to ovule in ovary, taking sperm with it
double fertilization
- one sperm nucleus fuses with egg nucleus –> 2n zygote
2. other fuses with the polar nuclei –> triploid endosperm
endosperm
unique to angiosperms, takes nutrients from parent gametophyte, repeatedly undergoes mitosis to produce starchy food source for embryo which is maintained as starch or taken up by embryo and packed into cotyledons
angiosperms appear
140 MYA, rapid diversification, viewed as “abominable mystery”, hypothesis: might have evolved from gymnosperm-like ancestors
angiosperm-like pollination in gymnosperms
non-conifer gymnosperms pollinated by insects or insects and wind, weevils develop in male cones of some cycads dusted with pollen, and adult weevils are attracted to sugary fluid produced by female cones, but fertilized zygote nourished by preexisting gametophyte cells
angiosperm-like fertilization in gymnosperms
several gnetophytes undergo something similar to double fertilization - one fertilized egg becomes zygote other aborts, still nourished by pre-existing tissues of gametophyte
2 reasons angiosperm fertilization more efficient
- food store (endosperm) develops only after fertilization
2. female gametophyte (embryo sac) reduced in size - inexpensive and quick to produce in only a few days
amborella trichopoda
oldest extant angiosperm, small dioecious shrub whose
flowers parts are in whorls of 5-13, greeny-yellow with undifferentiated petals and sepals, grows only in New Caledonia, wind and insect-pollinated
monocot classification
monophyletic 65 000 species
dicot classification
paraphyletic, but eudicots (165 000) monophyletic
monocot characteristics
1 cotyledon, parallel veins, scattered vascular tissue, fibrous roots, 1 opening in pollen, flower parts in 3s
eudicot characteristics
2 cotyledons, netlike veins, ringlike vascular tissue, taproot, 3 openings in pollen, flower parts in 4s or 5s
angiosperm importance to humans
80% of calories from 6 species, crops grown for oil, spices + flavour, sugar, drugs
teosinte
cob with less kernels selectively bred to become domestic corn
transgenic golden rice
contains genes from daffodils that make betacarotene (Vitamin A)