Plant responses Flashcards
What are tropisms?
Directional growth in responses to environmental cues such as light and gravity
Limitations on plants
They are rooted and not mobile without a rapidly responding nervous system - they are coordinated organisms that show clear responses to their environment, communication between cells and even between different plants
Timescale of plant responses?
Much slower than animal responses
Hormones
Chemicals that are produced in one region of the plant and transported both through the transport tissues and from cell to cell and have an effect in another part of the plant
Important plant hormones?
Auxins, gibberellins, ABA and ethene
Auxins role
Control cell elongation, prevent abscission, maintain apical dominance, stimulate release of ethene, tropisms, involved in fruit ripening
What is abscission?
Leaf fall
Gibberellins
Causes stem elongation, triggers mobilisation of food at germination, stimulate pollen tube growth in fertilisation
Ethene
Causes fruit ripening and promotes abscission in deciduous trees
ABA
Maintains dormancy of seeds and buds, stimulates cold protective responses (antifreeze production), stimulates stomatal closing
How do chemicals interact?
Not in isolation - they work at low concentrations so isolating and measuring them is not easy ; there are multiple interactions
Plant hormones and seed germination
When the seed absorbs water, the embryo is activated and begins to produce gibberellins which stimulate the production of enzymes that break down the food stores found in the seed
Embryo uses these food stores to produce ATP for building materials so it can grow and break out through the seed coat
Giberellins switch on genes which code for amylases and proteases - digestive enzymes required for germination
ABA reacts antagonistically with Gibberellins and that it is the relative levels of both hormones which determine when a seed will germinate
Where is food store in dicot seeds?
Cotyledons
Where is food store in monocot seeds?
Endosperm
Experimental evidence supporting the role of Gibberellins in germination
Mutant varieties of seeds have been bred which lack the gene that enables them to make gibberellins - these seeds do not germinate
If gibberellins are applied to the seeds externally
They then germinate normally
If giberellin biosynthesis inhibitors are applied?
They do not germinate as they cannot make the gibberellins needed for them to break dormancy - once removed, they then germinate
Auxins
Growth stimulants produced in plants - small quantities have powerful effects and they are made in cells at the tip of the roots and shoots and in the meristems ; move down the stem and up the root both in the transport tissue and from cell to cell
Auxins effect on plant growth 1
Stimulate the growth of the main apical shoot by affecting the plasticity of the cell wall ; presence of auxins means the cell wall stretches more easily
How does auxin cause cell wall to stretch?
They bind to receptor sites in the cell membrane, causing a fall in pH to about 5 ; this is the optimum pH for the enzymes needed to keep the walls very flexible and plastic
As the cells mature, auxin is destroyed and as the hormone levels fall, the pH rises so the enzymes maintaining plasticity become inactive (wall then becomes rigid and more fixed in shape and size)
Example of an auxin
IAA (indoleacetic acid)
As levels of auxin increase?
pH decreases
High concentrations of auxins
Suppress growth of lateral shoots - results in apical dominance ; lateral shoots are inhibited by the hormone that moves back down the stem so they do not grow very well (unlike the growth in the main shoot) - further down the stem the auxin concentration is lower so later shoots grow more strongly
If apical shoot is removed, auxin producing cells are removed and so there is no auxin and so as a result the lateral shoots grow faster (freed from apical dominance)
If auxin is applied artificially to the cut apical shoot
Apical dominance is reasserted and lateral shoot growth is suppressed
Low concentrations of auxin
Promotes root growth - up to a given concentration, the more auxin that reaches the roots the more they grow - auxin is produced by the root tips and auxin also reaches the roots in low concentrations from the growing shoots
If the apical shoot is removed, then the amount of auxin reaching the roots is greatly reduced and root growth slows and stops - replacing the auxin artificially at the cut apical shoot restores the growth of the roots
High auxin concentrations
Inhibit root growth
Gibberellins role
Affect the length of the internodes - regions between the leaves on a stem
Variations in plant size and hormones?
Plants that have short stems produce few/no gibberellins; dwarf plants - without gibberellins, shorter stems, reduces waste and makes the plant less vulnerable to damage by weather and harvesting
What should be done at the end of an experiment on plant hormones?
Important to make serial dilutions to observe the effects of different concentrations of the hormones as they can have different effects on growth - usually involve large numbers of plants, spread of data from each experimental group should be measured using standard deviation
Synergism
Different hormones working together, complementing each other and giving a greater response than they would on their own
Antagonism
Substances have opposite effects - one promoting growth and one inhibiting it, balance will determine the response of the plant
Abiotic stresses
Changes in day length, cold and heat, lack of water, excess water, high winds and changes in salinity - plant responses involve both physical and physiological adaptations
Lack of water adaptations
Thick cuticles, hairy leaves, sunken stomata or a wilting response in hot, dry or extremely windy conditions - develop aerenchyma if they grow in an aquatic environment
What causes plants to change in physical appearance?
Range of daylight hours and temperatures vary as well - thus they affect the rate of photosynthesis so seasonal changes have a big impact on the amount of photosynthesis possible
What point then comes after physical changes?
Amount of glucose required for respiration to maintain the leaves and to produce chemicals from chlorophyll that might protect them against freezing is greater than the amount of glucose produced by photosynthesis - a tree that is in leaf is more likely to be damaged or blown away by winter gales which means deciduous trees in temperature climates lose all of their leaves in winter and remain dormant until the days lengthen and temperatures rise again in spring
Daylight sensitive
Lack of light that is the trigger for change - this is known as photoperiodism ; breaking of the dormancy of lead buds so they open up and time of flowering in a plant
Sensitivity of plants to day length?
Results from a light-sensitive pigment called phytochrome - exists in two forms Pr and Pfr ; each absorb a different type of light and the ratio of Pr to Pfr changes depending on the levels of light
Abscission?
Lengthening it the dark triggers a period of dormancy during winter months and leaf fall
Falling light = falling concentrations of auxin ; leaves respond to the falling auxin concentrations by producing the gaseous hormone ethene - at the base of the leaf stalk is a region called the abscission zone, made up of two layers of cells sensitive to ethene
Ethene turns on genes - producing new enzymes which digest and weaken the cell walls in the outer layer of the abscission zone (SEPARATION LAYER)
What else happens building up to leaf fall?
Vascular bundles which carry materials into leaf are sealed off and fatty material is deposited in the cells on the stem side to form a protective scar when the leaf falls (preventing entry of pathogens) ; cells deep in separation zone respond to hormonal cues by retaining water and swelling, thus more strain on outer layer
ABIOTIC STRESSES like low temperatures or strong winds finish the process and the strain is too much, leaf separates from the plant leaving a neat scar behind
Preventing freezing
If cells freeze, membranes disrupted and thus they will die - but plants have evolved mechanisms that protect their cells in freezing conditions ; cytoplasm of plant cells and sap in vacuoles has solutes which lowers the freezing point and some plants produce sugars/polysaccharides/amino acids/proteins which act as antifreeze to prevent cytoplasm from freezing or protect the cells from damage even if they do freeze