motor control 1 Flashcards
what does motor control involve
dynamically changing mix of conscious and unconscious regulation of muscle force, informed by continuous and complex sensory feedback, operating in a framework sculpted by evolutionary pressures
types of motor control
- Voluntary: walking, talking ect
- Goal-directed: conscious, explicit, controlled
- Habit: unconscious, implicit, automatic
- Involuntary: eye movements, facial expressions, postural muscles, diaphragm ect
sensorimotor system key concepts
- Motor control governed by lower and upper motor neurons.
- The lower motor neuron begins (has its cell body) in brainstem or spinal cord and projects to the muscle
- The upper motor neurons originate in higher centres and project down to meet the lower motor neurons
muscle and muscle fibre activation
Individual muscle fibres act in an ‘all-or-none’ manner, and so control of muscle force depends on the way in which lower motor neurons activate different types of muscle fibre
antagonistic arrangement - combined co-ordinated action
skeletal muscle - muscle contraction
- A skeletal muscle comprises several muscle fasciculi (group of muscle fibres)
- A muscle fasciculus comprises several muscle fibres (= muscle cells)
- A muscle fibre is constituted of several myofibrils
- Myofibrils contain protein filaments: Actin and Myosin myofilaments
- When the muscle fibre is depolarised actin and myosin slide against each other which produce muscle contraction
- When a muscle contraction occurs a nerve impulse is sent to the neuromuscular junction that causes the release of acetylcholine – causes biochemical cascade.
rigor mortis
- ATP (provides energy for cells) is required to break the bond between the myosin head and the actin filament
- ATP is produced by oxidative metabolism, which stops upon death
- So the muscle become contracted and remain that way until enzymes begin to disrupt the actin/myosin
motor unit
- Motor unit = single alpha (α) motor neuron + all the extrafusal muscle fibres it innervates
- Different motor neurones innervate different numbers of muscle fibres – fewer fibres means greater movement resolution - those innervating finger tips and tongue
- The motor unit is the final common pathway for motor control
- More motor units fire – more fibres contract – more power
size principle
- Units are recruited in order of size (smallest first)
- Fine control typically required at lower forces
- E.g. eye = 3 motor units, trunk = over 1000 units
fast and slow muscle fibres
- Slow – e.g. posture
- Fast fatigue resistant – e.g. running/walking
- Fast fatigable – e.g. jumping
- Training and exercise lead to changes in the thickness of muscle fibres and the proportion of different muscle fibre types
the motor pool
- All the lower motor neurons that innervate single muscle
- The motor pool contains both the alpha and gamma motor neurons
- Motor pools are often arranged in a rod like shape within the ventral horn of the spinal column
cell bodies in the ventral horn activated by:
- Sensory information from muscle
* Descending information from brain
sensing in muscles - key part of proprioception
Muscles can be contracted or relaxed to provide movement, but a good control system (the CNS) needs to know two things:
- how much tension is on the muscle; - Golgi tendon organs sense tension
- what is the length (stretch) of the muscle – muscle spindles sense stretch
Golgi tendon organs – muscle tension (force)
- The GTO is within the tendon (where the muscle joins to bone)
- Mostly, it sends ascending sensory information to the brain via the spinal cord about how much force there is in the muscle
- Critical for proprioception
- Under conditions of extreme tension, it is possible that GTOs act to inhibit muscle fibres (via a circuit in the spinal cord) to prevent damage
reflexes
• Stretch detected – travels along muscle spindle – sensory neuron – synapses – signal sent across alpha motor neurons to extrafusal muscle fibres
intrafusal fibres are innervated separately, by gamma motor neurons
• They keep the intrafusal fibres set at a length that optimises muscle stretch detection
withdrawal reflex
- Reciprocal innervation of antagonistic muscles explains why the contraction of one muscle induces the relaxation of the other
- Permits the execution of smooth movements