V5 Flashcards
Structural features of the vegetation:
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Vegetation can be described by different characteristics:
- height of vegetation cover
- density of vegetation cover
- root penetration depth / intensity
- species composition !!!
- spectrum of life forms !!!
Structural features of the vegetation:
- in the course of evolution, a large number of species have evolved in adaptation to very different site conditions
- but at the same time only a small number of life and growth forms have developed
â âgenotypic differences + similar appearance and function in the ecosystem (= ecological equivalence)
Structural features of the vegetation:
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Description of a plant stand by:
- species composition > plant communities
- spectrum of life forms > plant formations
â Plant formations give information about abiotic environmental conditions by their shape = physiognomicâecological vegetation units
â different classification systems
Life and growth forms:
- includes total habitus (size, shape, outline) and life span
- plants: especially the shape of the leaves, the type of branching, the degree of lignification or the location of the renewal buds
- animals: additionally the predominant nutrition, the degree of the mobility or prominent behaviors
- Life forms after RAUNKIAER
Life forms after Raunkiaer:
Ecosystem characteristics:
- Ecosystem turnover
- Growth or defense (primary vs. secondary metabolism)
- Plant interactions (competition, mutualism, niches, resilience, valence)
Turnover:
- Net primary production (NPP) depending on site conditions:
- duration of the vegetation period
- global radiation
- air temperature during vegetation period
- water availability during vegetation period
- availability of mineral nutrients
- â 4. climatic
-
further turnovers:
- animal feed (=secondary production)
- Litter decomposition
- Humification
- Mineralization
- Net ecosystem exchange (NEE)
-
Ecosystem respiration:
- Reco = CO2 release from soil + plant
â NEE = GPP â Reco
Change in primary production, biomass growth, litter and respiration:
- Ecosystems are dynamic, longest dwell time in the mature phase
Model scheme of a natural/nearânatural ecosystem:
Primary production:
photosynthesis and respiration
- light can be the limiting factor
Phytomass and primary production in forest systems:
- esp. assimilation areas (LAI)
Primary production and evapotranspiration:
- Water use efficiency (g formed dry substance / liter of transpired water)
- Relation between production and water consumption
Secondary production (animal feed):
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Animals:
- heterotrophic creatures
- nutrition directly or indirectly from organic products of primary producers
- Consumers
- Secondary producers
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Biofages (live substance eaters):
- herbivores, carnivores, omnivores
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Dead substance eaters:
- detrivors ( â together with bacteria / fungi (saprobionic plants): decomposers)
-
quantitative importance of biophages:
- a few percent of the aboveâground phytomass eaten by herbivores
Four main groups of heterotrophic organisms:
Mineral turnover:
Primary and secondary metabolism:
- Tradeâoff between primary and secondary metabolism
- what is put into defense is no longer available for growth
- Secondary metabolites often âexpensiveâ
Plant defense:
- Response to stress caused by environmental conditions (pests, climate, air pollution, scarcity of resources, competition, âŚ)
- New production of secondary metabolites
- Supply of secondary metabolites
- âToxinsâ
- âWarning signalsâ
- âCalls for helpâ
- Triggering of numerous functions (e.g. maturation)
â not absolutely necessary for cell survival but useful or necessary for the whole organism
Plant defense, Example:
desert mugwort (WĂźstenâBeifuĂ) and wild tobacco
- in one group mugwort pruned, other group unchanged
- emission of VOCs
- tobacco near the pruned mugwort plants switches metabolism to defense
- significantly less feeding damage by locusts
- âeavesdroppingâ on neighboring plants / species
Interaction of individuals of the same or different species:
- Competition
- Mutualism (mutual benefit)
- Dependence on environmental conditions
â Result: siteâspecific species clusters
â Plant communities and formations
Ecological niche:
- The entity of environmental factors that a uniform group of plants or animals (e.g. species) needs for their survival
- biotic and abiotic factors combine to form a multidimensional hyperspace
- functional relationship between a species and the ecosystem
- within the niche space: competition, mutualism, symbioses
fundamental niche â realized niche
Competition:
- nteraction of organisms (of the same or different species) based on competition for limited resources
- intraâ and interspecific competition
- mindâexperiment: pure vs. mixed culture
-
Picture:
- Species 2 could also grow under extreme conditions, but is displaced by more competitive plants under these conditions
Physiological valence:
- every plant species can exist from its physiological constitution under wide ranges of environmental conditions
â physiological range of existence / physiological valence
- depending on the intensity of the influencing factor, different growth and distribution
- range of highest production output: physiological optimum
Ecological valence:
- in case of occurrence of several species: mutual displacement/enforcement from more or less large parts of the physiological area of existence â ecological subsistence area / ecological valence
- ecological optimum
- ecological and physiological optimum coincide only in a few cases (mostly at the physiological edge)
- misleading or wrong: âdrought lovingâ, âin need of warmthâ, âlime seekingâ
- species only more competitive â âdroughtâbearingâ
Effects of competition:
- intraspecific competition for water, nutrients and radiation
- e.g. same total biomass production with different numbers of individuals of the same species (e.g. three or six Lolium perenne and three Daucus carota plants)
- die off of the âweakâ individuals â Selection of the best adapted individuals â Optimization of the genetic adaptation of the species
- interspecific competition
- e.g. three Lolium and three Daucus plants
- Lolium biomass almost as in pure culture
- Daucus biomass only at about 10% of the possible biomass
- total biomass (Lolium + Daucus) same as before
-
with a threeâweek growth advantage of Daucus:
- Lolium only 40%
-
at threeâweek growth advance of Lolium:
- Daucus practically at 0
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Adaptation to competition:
- little energy in generative apparatus, more in height growth
-
e.g. with Daucus ahead:
- Lolium compared to normal growth 40% stalk and leaf biomass, only 20% ear biomass (few seeds) â less offspring â extinction after some generations
- Competition for nutrients and water underground, light above ground
â Plant individuals recognize competing individuals very early via their photoreceptors (e.g. by the red/far red ratio)
Mutualism (mutualistic symbiosis):
- Interrelation between living beings of two species
- at least one partner benefits compared to living alone, the second partner is at least not affected
- Increase of the competitive strength
- Colonization of more extreme habitats
- Improvement of absorption of water and minerals
- Making nutrients available from deeper layers
- Water storage in cushions
â positive interaction
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Examples:
- Central European forest trees and mycorrhiza
- organization form of lichens (algae + fungi) â â196°C to +70°C
Plant interactions:
- each plant individual is involved in a permanent network of relationships between all stand individuals
- competition is essential relationship
- but also mutual support
Influence of plants on environmental variables:
- plants influence existential requirements of other species but also of the own species (rejuvenation)
-
light / radiation:
- strong reduction â strong selection of species due to lack of light
-
precipitation:
- interception, fog scavenging â increase, decrease or redistribution in the stand
-
soil water content:
- by evapotranspiration lowering of the groundwater level â drying of the upper soil layers
- soil nutrient content:
- conifers â acid tolerant plants are grown; Robinia â N cycle
Ecogram:
Ecosystem characteristics:
dynamic equilibrium
- quasi-stationary state of an ecosystem during an observation period (inputs and outputs of energy and matter in equilibrium)