Phloem translocation Flashcards
phloem transports products of photosynthesis from
mature leaves to areas of growth and storage
phloem conducts – molecules
signal
phloem – water
redistributes
phloem extends throughout the plant body and is – with xylem
parallel
in trees with secondary growth, the phloem is on the outside of the – and only the inner layer of phloem is functional as outer is crushed by xylem
vascular cambium
phloem is removed by removing bark
girdling
girdling tree – ceases between root and leaves
sugar transport
sieve tube elements
angiosperms
sieve cells
gymnosperm
store and release food molecules
parenchyma cells
protection and strengthen tissue
fibers and sclereids
sometimes – (latex containing) for protection
laticifers
only – are directly involved in translocation
sieve elements
small veins of leaves and primary vascular bundles of stems are typically surrounded by a – which are compactly arranged colorless cells isolating the veins from the intercellular spaces of the leaf
bundle sheath
mature sieve elements contain
mitochondria, plastids, smooth ER
sieve element’s cell walls are – and thus remain soft and flexible
non-lignified
where pores connect to the other sieve elements
sieve areas
pore range from
less than 1 to 15 micro m
in angiosperms, sieve areas on the end walls can differentiate into – with larger pores and
sieve plates
sieve tube elements can connect to form –
sieve tubes
in gymnosperms, – cover the sieve areas
smooth ER
dive cells contain sugars and organic molecules in a fluid solution
sap
sieve tube elements also contain – which are tubular, fibrillar, granular, or crystalline
P proteins
P proteins seal damaged sieve elements by plugging off –
sieve plate pores
relatively few plasmodesmatal connections to cells except its own sieve element; symplastically isolated from surrounding cells
ordinary companion cells
finger-like wall ingrowths of cell walls facing away from the sieve tube elment
transfer companion cells
many plasmodemsata connections with bundle sheath cells, well-suited for taking up solutes via cytoplasmic connections
intermediary companion cells
ordinary companion cells – loading
apoplastic
transfer companion cells – loading
apoplastic
intermediary companion cells – loading
symplastic
sources are typically mature leaves; others include storage organs –
storage root of beets
sinks = – organs including roots, developing flowers and immature leaves
nonphotosynthetic
phloem sap can be collected using – natural syringes
aphid stylets
most abundant substance in phloem
water
sugar in phloem are –
nonreducing
typical range of phloem sap
30 cm to 1.5 m per h
pressure gradient of bulk flow is established by
loading and unloading of sugars into sieve tubes
sieve plate – slows equilibration allowing a standing pressure gradient to exist and to drive flow
resistance
sucrose moves from producing mesophyll cells to cells adjacent to the sieve elements in the leaf minor veins, only a few cell diameters
short distance transport pathway = symplastic
sugars are transported into the sieve elements and companion cells
phloem loading
sugars are translocated away form the source via
long-distance transport
– loading requires energy
apoplastic
H+-ATPase transports H+ into the apoplast and a – transport sucrose form the apoplast into the sieve element-companion cell complex
sucrrse H+ symporter
explains how different solute concentrations can be generated in different adjacent cells connected by plasmodesmata
polymer-trapping model
sucrose synthesized in the mesophyll diffuses form the bundle sheath cells into the – via small plasmodesmata
intermediary cells
In the intermediary cells, – and – are syntehsized
raffinose and stachyose
larger raffinose and stachyose diffuse through the larger plasmodesmata to the –
sieve element
– loading can also occur given high enough sugar production rates
passive
in dev leaves, first whole leaf is a sink and sugars are unloaded from the – in the unexpanded leaf
major veins
as leaf matures, major veins spaced apart and – dev between them
minor veins
when unloading ceases and loading begins only in –
minor veins
signal molecules are – from sources
loaded
GFP expressed in – cells to show loading and unloading
companion cells