Introduction to Comparative Vertebrate Anatomy Flashcards
Studies the descriptive and functional morphology of vertebrates
Descriptive - structure of vertebrates
Functional - significance of the structure
Comparative Anatomy
A study of evolution to phylogeny
It can be considered as vertebrate evolution
Comparative Anatomy
study of structural evolution
Morphology
study of description of structures
Anatomy
Tail appearing outwardly symmetrical but with the backbone passing into the upper lobe, as in all higher fish
homocercal
Tailhaving unequal upper and lower lobes, usually with the vertebral column passing into the upper.
heterocercal
Big Five in chordates
post anal tail
dorsal hollow nerve cord
pharynx
notchord
endostyle
No vertebrate will proceed without passing through these five characteristics in embryonic development
true
_______ have been grouped into a single taxon
Protochordates and chordates
is a rod of living cells ventral to the CNS and dorsal to the alimentary canal
provides skeletal support throughout most of the length of a chordate
Notochord
most vertebrates, a more complex, jointed skeleton develops, and the adult retains only remnants of the
embryonic notochord
Fate during development of Notochord and Vertebral Column in head region
incorporated into skull floor
Fate during development of Notochord and Vertebral Column in Trunk and tail
surrounded by cartilaginous or bony vertebrae
Fate in adulthood development of Notochord and Vertebral Column in Trunk and tail in Protochordates
chief axial skeleton surrounded by notochord sheath
Fate in adulthood development of Notochord and Vertebral Column in Trunk and tail in Agnathans
similar to protochordates with addition of lateral neural cartilages
Fate in adulthood development of Notochord and Vertebral Column in Trunk and tail in Fishes and amphibians
persist the length of the trunk and tail within the centrum
Fate in adulthood development of Notochord and Vertebral Column in Trunk and tail in Reptiles, birds and mammals
disappears and becomes pulpy nucleus in mammal vertebrae
Vertebrae parts
Centrum (deposited around notochord)
Neural arch (forms over spinal cord)
Various processes
develops from a plate of ectoderm that rolls into a tube dorsal to the notochord
Formed by the process of invagination
Dorsal Hollow Nerve Cord
The nerve cord develops into the central nervous system:
Brain
Spinal cord
Common to hemichordates and chordates
can be perforated by openings (slits) to either the exterior or an atrium (reflecting common ancestry)
Membranous digestive and respiratory organ located at the back of the mouth that serves as passageway of food and air
Pharynx
The ______ Vital part of craniate embryo that produces:
Gills of fishes (permanent slits)
Lungs of tetrapods (temporary slits)
Skeleton & musculature of jaws
Endocrine glands that regulate metabolic rates in all body cells & maintain appropriate calcium levels in bones, other tissues & circulating blood (parafollicular cells & parathyroid glands)
Gives rise to middle ear cavity of tetrapods (auditory and tympanic cavity)
Provides initial cells of immune system during fetal life & shortly after (thymus in humans)
Pharynx
A column of tissue which separates each embryonic pharyngeal pouch or slit from the next
Pharyngeal Arches
Pharyngeal Arch typically has 4 components or blastemas from which these components develop:
Skeleton
Muscles
Nerves
Blood vessels
Total of _________ of Pharyngeal arches develop in cranio-caudal sequence
6 pairs
is complex, and contributes to both upper and lower jaw
First arch
Arch _
Skeletal Elements
Meckel’s cartilage
Malleus
Incus
Muscles
Muscles of mastication
Anterior belly of digastricus
Tensor tympani
Tensor palati
1
Arch 1
Cranial Nerve
Trigeminal nerve (mandibular division)
Arch _
Skeletal Elements
Stapes
Styloid process
Upper part of hyoid
Muscles
Stapedius
Stylohyoid
Muscles of facial expression
Posterior belly of digastricus
2
Arch 2
Cranial Nerve
Facial nerve
Arch _
Skeletal Elements
Lower part of hyoid
Muscles
Stylopharyngeus
3
Arch 3
Cranial Nerve
Glossopharyngeal nerve
Arch _
Skeletal Elements
Laryngeal cartilages
Muscles
Cricotyhroideus
Pharyngeal constrictors
Instrinsic laryngeal muscles
4 - 6
Arch 4 - 6
Cranial Nerve
Vagus Nerve
Supportive skeletal elements (pharyngeal skeleton)
Pharyngeal Arches
Pharyngeal Arches have ___________ (branches 5, 7, 9 and 10) which innervate muscles and provide sensory input to the brain
Cranial nerves
Pharyngeal Arches have an __________ that connects the ventral and dorsal aortas
aortic arch
Striated muscles that operate the Pharyngeal Arches
branchiomeric muscles
Ectodermally-lined grooves on the outside of the embryonic pharynx
Pharyngeal Clefts
Supportive skeletal elements (pharyngeal skeleton)
Striated muscles that operate the arch (branchiomeric muscles)
Cranial nerves (branches 5, 7, 9 and 10) which innervate muscles and provide sensory input to the brain
An aortic arch that connects the ventral and dorsal aortas
Pharyngeal Arches Components
Ectodermally-lined grooves on the outside of the embryonic pharynx
are lost in development because of proliferation of the underlying mesoderm
Pharyngeal Clefts
an important cleft because this Pharyngeal Cleft it develops into the external auditory meatus of the ear and provides the outer epithelium of the tympanic membrane
first cleft
Arises as diverticula of endoderm or foregut
Grows toward the surface of the animal
Establish the limits of the pharynx
Contribute to the development of a surprising diversity of structures
Pharyngeal Pouches
Maximum number in jawed craniates in basal shark and living agnathans
8 - basal shark
15 - living agnathans
grows toward each Pharyngeal pouch and is separated from each other via the pharyngeal plate
Pharyngeal clefts
Pharyngeal Pouch _ Derivatives
Middle ear cavity
Endodermal aspect of tympanic membrane
Pharyngotympanic tube
1
Pharyngeal Pouch _ Derivatives
Palatine tonsil
2
Pharyngeal Pouch _ Derivatives
Inferior parathyroid gland
Thymus
3
Pharyngeal Pouches __ Derivatives
Superior parathyroid gland
Parafollicular cells of the thyroid gland
4-5
are formed between the pharynx and the exterior when the pharyngeal plate ruptures
Pharyngeal Slits
Pharyngeal Slits may open to the exterior as slits
True
where Permanent Pharyngeal slits are
adults that live in water and breathe via gills
where Temporary Pharyngeal slits are
adults live on land
is a glandular groove in the floor of the pharynx and is involved in filter feeding
Endostyle
is an endocrine gland that produces hormones
Thyroid gland
Both arises from the floor of the pharynx and are involved in iodine metabolism
Thyroid gland and Endostyle
Chordates have a ________ posterior to the anus
contains skeletal elements and muscles
Post-Anal Tail
In many species, the tail is greatly reduced during
embryonic development
It provides propelling force in many aquatic species
Post-Anal Tail
origin of protostomes
coelom originates from split of the mesoderm, forms the mouth
origin of deuterostomes
coelom originates from outpouching of the gut, forms the anus
are animals that have a notochord in the embryo stage at least
Chordates
are chordates with a neurocranium or braincase
Craniates
chordates with vertebrae
Vertebrates
appear during embryonic development after the notochord has formed
Vertebrae
craniate body consists of three regional components:
Head
Trunk
Postanal tail
Contains special sense organs for monitoring the external environment - eyes, ears, nose
Head
is at least large enough (cephalization) to receive and process essential information and to provide appropriate stimuli to the body musculature
Brain of chordates
in some species for acquiring, retaining and macerating food
Jaws
for respiration in fishes
Gills
Centralization or localization of nervous structures and functions in the head with accompanying dominance of the head
Cephalization
More and more cephalization marked the higher one ascends in the animal kingdom
true
Particularly prominent as a structural and functional feature of the vertebrates
Cephalization
Contains the coelom which houses most of the visceral organs
Trunk
surrounds the coelom and consists chiefly of muscle, vertebral column and the ribs
body wall
the trunk is Associated with
paired pectoral and pelvic appendages
which is a narrow extension of the trunk connects the trunk with the head
Neck
Commences at the anus or vent
Consists almost exclusively or a caudal continuation of body wall muscles, axial skeleton, nerves and blood vessels
Postanal Tail
Postanal Tail is Present in all embryos, though adult form may lose the tail along development
true
position when a quaruped stands erect on all four limbs, facing left, with the tail slightly raised
Anatomic Position
any surface, real or imaginary, along which any two points can be connected by a straight line
Anatomic Plane
divides the head, body or any limb longitudinally into equal right and left halves
Median Plane
There is only one single median plane
true
Median and medial are not synonymous
true
Median is a
Medial is a
plane
directional term
passes through the right or left side of the body parallel to the median plane
Sagittal Plane
passes through the head, body, limb or organ at right angles to the structure’s long axis or the median plane
Transverse Plane
runs at right angles to both the median and transverse planes (horizontally) dividing the body into dorsal and ventral potions
Dorsal Plane
Occurs in an organism if it can be cut into two identical halves through any cut that runs through the organism’s center
Spherical
These organisms resemble a pie where several cuttingplanesproduce roughly identical pieces
They have a top and a bottom (dorsal and ventralsurface) only
Radial organisms
An organism with ___________ symmetry exhibits no left or right sides
radial
Combination of radial and bilateral symmetry
Biradial
Only one plane, thesagittal plane, will divide an organism into roughlymirror imagehalves (with respect to external appearance only)
Bilateral
Internally, only the organ systems are not bilaterally symmetrical
digestive and lymphatic system
Regular repetition of body parts along the cranio-caudal axis
Metamerism or Segmentation
Body of segmented animals is composed of longitudinal series of divisions in each of which all or most of the body systems are represented
metamere, segment or somite
No external evidence of segmentation in most adult reptiles, birds, & mammals
But internally a series of muscle segments are visible
true
Serial arrangement of vertebrae, ribs, spinal nerves, embryonic kidney tubules, segmental arteries & veins)
true
Each segment may or may not be marked externally by a constriction of the body wall
Metamerism or Segmentation
segmentation of the animal body into nearly like segments
Homonomous segmentation
various segments differ from each other
Heteronomous segmentation
Cranium
Three-part brain
Neural crest and its derivatives
Paired external sense organs
Cartilage
Craniate
Vertebral column
Two semicircular canals
Electroreception
Lateral line system with multicellular neuromasts
Number of soft tissue specializations
Vertebrate Characteristics
Integument - epidermis and dermis
Respiratory mechanism - external
Coelom
Other Craniate Characteristics
Paired external sense organs
olfactory, optic and otic with a single semicircular canal, lateral line system with unicellular sense organs
epidermis and dermis of craniates
Integument
coeloms in Craniates
Pericardial cavity - heart
Pleural cavity - lungs
Peritoneal cavity - abdominal organs
Scrotal cavity - testes
Digestive system - cloaca or anus
Urogenital organs
Circulatory system - single or double-circuit
Skeleton - axial and appendicular
Muscles - skeletal, smooth and cardiac
Sense organs - exteroceptors, proprioceptors and visceral receptors
Other Craniate Characteristics