FINAL Flashcards
what does vertebrate translate to
animal with metameric skeletal elements supporting a dorsal hollow nerve cord
what does anatomy translate to
anatomia, structure, morphology
what is metamerism
repeating units
what is one flaw in the word vertebrate
hagfish dont have vertebrae
what does anatomia translate to
cutting up or dissection
what is interspecific
between species
what is intraspecific
within species
give an example of interspecific
locomotion and bipedalism of humans versus gorillas
give an example of intraspecific
the different kinds of vertebrae in a human
what is the inominate bone
ilium + ischium + pubis
what are the three subphyla of the chordates
cephalochordata, urochordata, vertebrata
what is found within the urochordata
tunicates (sea squirts)
what is found within the cephalochordata
amphioxious (lancelets)
a vertebrate is a deuterostome, what does this mean
it refers to the anus being formed first from the blastopore during development
what is an example of a hemichordate
acord worm
what is osteology
the study of bones
what is a craniate
an organism that possesses a cranium (bone or cartilage)
what are the three classes in subphylum urochordata
ascidiacea, larvaceae, thaliaceae
what subphylum has their notochord only in the tail
urochordates
what subphylum has notocord in head and tail
cephalochordates
what are the 5 chordate features
notochord, dorsal hollow nerve chord, pharyngeal openings, post anal tail, endostyle
what is a notochord
endoskeletal structure
what is the dorsal hollow nerve chord
main nerve chord of the body, anteriorily forms the brain
what is the function of the post anal tail
generally used for locomotion or balance
what is the function of the endostyle
it acts as a food trap in some organisms (thyroid in some)
what does deuterostome translate to
secondary mouth
what are the deuterostomes
chordates, echinoderms, hemichordates
how is short term ATP synthesis done in deuterostomes
phosphocreatine
how is short term ATP synthesis done in non-deuterostomes
phosphoargine
whats the difference between phosphocreatine and phosphoargine
same function, different molecule
why would an organism use phosphocreatine or phosphoargine
gives energy faster than anaerobic respiration
what was the first theory of chordate origins and who did it come from
annelid-arthropod
dorsal ventral inversion
1822 geoffroy st. hillaire
what was the second theory of chordate origins and did it come from
auricularian hypothesis
chordates arise from echinoderm larvae and tunicate larvae
1928 Garstang
what is the current theory of chordate origins
earliest chordates came from cephanochordates which probably came from hemichordates
give an example of species that would fit into the current theory of chordate origins
acorn worm –> lancelet –> cat
what is an auricularian in short
type of deuterostome larvae that is a dipleureae (comes from sea cucumbers)
why was st hillaire’s chordate origins theory proven wrong
convergent evolution
what is histology
the study of tissues
what is a tissue
a group of similar cells performing a common function
what are the ways cells can be performing a common function
appearance, morphology, embryonic
what is a cell in relation to histology
fundamental living elements
what is the matrix in relation to histology
acellular, non-living components surrounding the cells (goop between the cells)
what are the four primary tissue types
epithelia, connective tissue, muscle, nervous tissue
what is epithelial tissue
the tissue that lines organs (inner most and outermost tissue)
describe epithelial tissue
absorbent, secretory, or protective
how are epithelial cell tissues classified
cell shape, stratification
what is the difference between apical and basal epithelial tissues
apical is top or surface cells, basal is bottom cells or inner cells where other cells adhere to
what is the density of epithelial cells
high cell density, little matrix
what does it mean that a tissue is stratified
it has layers
what are the three types of epithelial cells
squamos, cuboidal, columnar
where can you find squamos cells (epithelial tissue)
kidney, lungs, lining of heart, blood vessels, lymphatic vessels
what is the function of cuboidal epithelium
secretion and absorption
where can cuboidal epithelia cells be found
kidney tubules, ducts, secretory portions of small glands, ovary surface
what is a pseudocolumnar epithelium
varying lengths of epithelial cells make it look like theres layers
what are connective tissues
lots of matrix (bone and cartilage)
how do epithelial cells adhere to basal membrane
by proteins
what do echinoderm larvae look like
bilaterally symmetrical, 2 bands cilia, complete gut
give example of auricularian theory
echinoderm to hemichordate to chordate
what are the periods in the paleozoic era
cambrian, ordovician, silurian, devonian, carboniferous, permian (can older students develop crappy personalities)
what are the periods in the mesozoic era
triassic, jurassic, cretaceous
when was the age of fishes
silurian, devonian, carboniferous, permian
what MYA was paleozoic era
575-230
what MYA was mesozoic era
230-65
how many MYA was pikaia
530
when did the gnathosomes appear
ordovician
when did amphibians appear
devonian
when did reptiles appear
carboniferous
when did therapsids appear
permian
when did mammals and dinosaurs appear
triassic
what were the ostracoderms
early agnathan fishes with dermal bone
what are the oldest bones found in fossil records
dermal bone
what are examples of dermal bone in modern day
turtle shells, fish scales
what is paedomorphosis
retention of juvenile characteristics in adults
what was myllokunmingia
lower cambrian craniate
what was haikouichthys
lower cambrian vertebrate
what are the 10 characteristics of a vertebrate
5 of chordate, integument with dermis and epidermis, complete digestive system, advanced circulatory system, excretory system, endocrine system
where is the vertebrate digestive system in relation to the spinal column
digestive system is ventral in relation to spinal column
what is the endocrine system
series of ductless glands with secretory products (hormones) released directly into the blood
why is it difficult to identify notochord cells
they look like adipose cells
what is the notochord replaced by in most vertebrates
usually replaced by vertebrae
what does the dorsal hollow nerve chord usually contain
usually has an anterior cranial expansion
what kind of expansion is a traditional brain
tripartite meaning three bulbs of expansion on the anterior end of the dorsal hollow nerve chord
what is typical of most vertebrates to do with their “brain”
encase it with a cranium (bone or cartilage)
what on the blastopore forms the notochord
the blastoporal lip
who experimented with putting two blastoporal lips on the same blastopore
spemann
what classes comprise the agnathans
ostracoderms, pteraspidomorphi, conodonita, myxinoidea, pteromyzontida
what does agnathan mean
absence of jaw (jawless fishes)
what is found in class chondrictes
cartilagenous fishes (sharks skates rays)
what is found in acanthodii
spiny fishes
what is found in actinopterygii
ray finned fishes
what is found in sarcopterygii
fleshy finned fishes
which portion of the skeleton is oldest
axial
what portion of the skelton is newest
appendicular
how many planes of sectioning are there to bilaterally symmetrical organisms
3 planes of symmetry
what are the three distinct origins of the mammalian skull
splanchnocranium, chrondrocranium, dermatocranium
which region of the skull is oldest
splanchnocranium
what is the splanchnocranium associated with
the pharynx
what is the chrondrocranium associated with
surrounds the bottom and the sides of the brain
what is the dermatocranium associated with
the top of the brain box
what is a demifacet
dent in thoracic ribs where rib touches
what is the first vertebrae
C1 is the atlas (holds up cranium)
what is the second vertebrae
C2 is the axis
what classes are in the teleostomi
acanthodii, actinopterygii, sarcopterygii
what class of fishes did mammals evolve from
sarcopterygii
if you are not an amniote then what are you
if not amniote then anamniote
what are the anamniotes
fishes and amphibians
what are the amniotes
birds, mammals, reptiles
what does it mean that youre an amniote
you dont have to deposit your eggs in water (can create own water in an egg)
bone and cartilage are very similar but primarily differ in what
they differ in their matrix
what are the three types of cartilage
fibrocartilage, elastic cartilage, hyaline cartilage
what are the two kinds of bone
spongy bone, compact bone
what are the caudata
salamanders
adipose cells are filled with fat, notochord cells are filled with what
filled with water
what does it mean to be a vacuoled cell
most of the cell is occupied by something else (ex. adipose and notochord)
with vacuoled cells there is usually a sheath covering the mass of cells. what is this sheath made of
collagen
what are the disks between vertebrae
annulus of fibrocartilage with nucleus pulpous in between
where can notochord cells be found in adult mammals
in the nucleous pulpous of the vertebral disks
what can mesenchyme produce
fibroblast, chondroblast, osteoblast, hematopoetic stem cell
what do fibroblasts produce
connective tissues (fat, skin)
what do chondroblasts produce
cartilage
what do osteoblasts form
bone
what do hematopoetic stem cells produce
blood
what is mesenchyme
embryonic tissue made of migrating cells thats not found in sheets, but found in blobs (pleuripotent cells (stem))
what does blast mean
forming
what is matrix composed of
collagen, elastic fibres, proteoglycans
what are proteoglycans
core protein with link proteins with glycosaminoglycans (looks like test tube cleaning brush)
what are the 6 types of glycosaminoglycans (GAG’s)
hyaluronate, chondroitin sulfate, heparin sulfate, heparin, dermatan sulfate, keratin sulfate
are GAG’s hydrophilic or hydrophobic
hydrophilic
Where is hyaluronate found (GAG)
synovial fluid, vitreous humour, ECM of loose connective tissue, large polymers, shock absorbing
where is chondroitin sulfate found (GAG)
cartilage, bone, heart valves
what is the most abundant of the GAG’s
chondroitin sulfate
where is heparan sulfate found (GAG)
basement membranes, components of cell surfaces
where is heparin found (GAG)
intracellular granules of mast cells, line arteries of lungs liver and skin
where is dermatan sulfate found (GAG)
skin, blood vessels, heart valves
where is keratin sulfate found (GAG)
cornea, bone, cartilage (makes things waterproof)
in general how is cartilage formed
mesenchyme to chondroblast then GAGs surround chondroblast then GAGs surrounded by a lacuna shell. once surrounded by lacuna its cartilage
whats a place you could find elastic cartilage
ears
whats the function of elastic cartilage
maintains shape of structure while allowing great flexibility (more elastic fibres in matrix)
where can fibrocartilage be found
intervertebral joints, pubic symphyses, discs of knee joints
whats the function of fibrocartilage
tensile strength with the ability to absorb compressive shock
what is the predominant fiber in fibrocartilage matrix
thick collagen fibers
where can hyaline cartilage be found
embryonic skeleton, covers ends of long bones, costal cartilage on ribs, cartilage of nose, trachea, larynx
whats the function of hyaline cartilage
supports and reinforces, resilient cushioning, resists comprehensive stress
what is osseous tissue
bone
what is compact bone
cortical bone thats outside bone (flat bone)
what is spongy bone
cancelous, inside spongy bone (network of osseous tissue)
what is osteogenesis
the formation of bone
what are the two types of osteogenesis
intramembranous
cartilage replacement
what is osteogenesis in the form of intramembranous bone formation
happens to flat bone of skull and clavicles, happens within fibrous connective membrane
what is osteogenesis in the form of cartilage replacement bone formation
happens in most skeletal elements, can be perichondral or endochondral
what is an ossification center
the site of bone formation
what engages osteogenesis
osteoblasts
what removes existing bone
osteoclasts
what maintains fully formed bone
osteocytes
in what kind of development is the first formative bone nonlamellar
endochondral and intramembranous
what is another name for nonlamellar bone
immature bone or woven bone
what is nonlamellar bone
lots of cells interspersed amongst irregularly strewn bundles of collagen
what is perichondral
on the outside
what is endochondral
on the innermost
what is lamellar bone
mature bone
what does osteogenesis start with
local aggregations of loosel arranged mesenchymal cells
in what kind of osteogenesis formation is cartilage an intermediate
endochondral
when someone stops growing what has happened
their epiphysial plates have closed
what are the three regions have a bone
epiphysis, metaphysis, diaphysis
what is another name for the metaphysis
epiphyseal plate
what is the first step of bone formation from endochronral
loose cellections of mesenchymal cells consense to form hyaline cartilage
what is the second step of bone formation in endochondral
bone collar forms in region of diaphysis
what deposits the bone collar
cells on inner surface of diaphysis become osteoblasts and deposit the collar
what is happening in the bone at the same time as the formation of the bone collar
inorganic calcium salts accumulate in the matrix to calcify cartilage in the core of the diaphysis
what is the function of calcium in bone formation
calcify cartilage, seal off chondrocytes
what is the function of sealing off chrondrocytes in bone formation
seal them off so they die so vasculature can invade
when is the primary center of ossification established
when osteoblasts appear in the core of the bone
what happens in the center of ossification
old bits of calcified cartilage bone become overlaid by new bone
what are trabeculae
transitional composites of new bone
when are trabeculae considered bone spiracles
when an ossified matrix predominates
what do osteoclasts signal in osteogenesis
signals active nature of bone remodeling through matrix deposition and removal
what is the difference between osteoblast and osteoclast
blast deposits, clast removes
where is the active area of cartilage growth, calcification, cartilage removal, and new bone deposition
epiphyseal plates
what do chondrocytes do in response to approaching ossification
chondrocytes proliferate and hypertrophy as surrounding matrix calcifies
what lengthens bone
proliferation of cartilage in the epiphyses
what contributes to an increase in bone girth
continued depositon of bone under diaphyseal periosteum
why can fish grow larger even if reached sexual maturity
only bird and mammals stop growing when they reach adult and sexual maturity
what is the basic steps of osteogenesis
cartilage calcifies, blood vessels invade epiphyses, osteoblasts appear, new bone deposited
when do secondary centers of ossification appear in humans
at 2-3 years of age
when does the zone of ossification take over cartilage proliferation
at or shortly after mammals reach sexual maturity
what are the 5 chordate characteristics that all vertebrates have
notochord, pharyngeal openings, post anal tail, endostyle, dorsal hollow nerve cord
what are the innovations of chordates that are characteristic to vertebrates
cranium and vertebrae
what are vertebrae
series of separate bones or cartilage blocks firmly joined together as a back bone that defines the major axis of the body
what are between vertebrae
intervertebral disks
what are the 3 basic parts to a vertebra
centrum, neural arch, hemal arch
what structural element did early vertebrates rely o
has vertebrae but still mainly relied on the notochord
where does the notochord remain in advanced vertebrates
in the nucleous pulpous in the intervertebral disks
what is the function of a cranium
supports sensory organs in the head and encases or partially encases the brain
what does cephalization mean
anterior clustering of specialized sensory organs
what are the three parts of the brain
forebrain, midbrain, hindbrain
what two kinds of cells give rise to most sensory organs in the head and are the source of most adult structures that distinuish vertebrates from chordates
neural crest cells, epidermal placodes
in what habitat are the earliest vertebrates found
marine waters
what are the three stages of vertebrate evolution
prevertebrates, agnathans, gnathostomes
what is a prevertebrate
suspension feeding organism that probably resembled amphioxious without burrowing habits
how did agnathans feed
use of a muscular pump to produce food bearing water current
where did the prevertebrates arise from
arose within protochordates
what two mechanical changes caused the shift from prevertebrate to vertebrate
pharynx developed encircling band of muscles, strong and springy cartilage replaced collagen in pharyngeal bars (this formed the muscular pump)
what removed the limits to size in early vertebrates
the loss of cilliary pump and the dominance of muscular pump
what did the muscular pump of prevertebrates allow for
allowed for larger body size, more complex feeding, addition of gills instead of pharyngeal openings
what are gills
complex, folded respiratory organs on pharyngeal pouches whose folds contain blood capillary beds capable of gas exchange
what are the folds of gills called
lamellae
what did the addition of gills allow for in vertebrates
allowed for increased respiratory demands made by organism due to increased activity
how did early vertebrates arise to the agnathan stage
appearance of muscular pharyngeal pump
what are the 5 agnathan groups
conodonts, pteraspidomorphi, ostracoderms, myxini, pteromyzontida
what kind of feeding did early gnathostomes posses
raptiorial and suction feeding
what removed size restriction of prey for gnathistomes
addition of jaws
if youre not a fish youre a ____
tetrapod
what are amniotes
vertebrates that wrap their embryos in an amnion
who are the anamniotes
fishes and amphibians
what are jaws derived from
phyrengeal arches
were ostracoderms vertebrates
no, notochord instead of vertebrae
ostracoderms were not considered craniates, why
they had a dermal bone head shield but not a chondral cranium
what do odontoblasts do
form dentin and enamel (mesenchymal derivatives)
how does dermal bone vary most significantly from other bone types
dermal bone has an enamel surface
what is the issue with having bone made of enamel and dentin
no blood vessels so its not reparable
what are the extant agnathans
hagfish and lampreys (myxini and pteryomyzontida)
what kind of larvae do lampreys have
ammocoete (kind of like amphioxious)
what kind of teeth do lampreys have
keratinized teeth (like finger nails)
lampreys and hagfish do not have arcualia when embryonic, what are arcualia
theyre embryonic structures that form vertebrae
whats the ecological difference between hagfish and lampreys
lampreys are ectoparasites
what kind of feeding do hagfish have
carrion feeding
what is carrion feeding
eat dead stuff
how many semicircular canals do lampreys have
2
how many semicircular canals do hagfish have
1
hagfish are unique in that they are isoosmotic, what is isoosmotic
organisms cells are in balance with the salt of the ocean
what organism displays knotting feeding when feeding
hagfish
what are the cyclostomes
hagfish and lamprey because they have circular mouths
where does the fossil data for hagfish and lampreys reach to
late devonian
all agnathans lack ____ and possess____
lack bone and possess a single nostril
what is the function of hagfish knotting
provides force to tear off food from a larger body
hagfish are hermaphrodites but they are not practicing hermaphrodites, what does this mean
they have both sets of gonads but only use one set, must have another hagfish present to reproduce
what kind of agnathan has vertebrae
some hagfish have embryonic vertebrae
what does it mean for water movement in the body of the hagfish that it is isoosmotic
no net flow o water in or out of the body (doesnt have to drink or pee)
what was most likely the direct ancestor to hagfish
invertebrates
how does a lamprey stay in place in moving water
grasps rock in mouth and holds it
what kind of feeding does the lamprey larva show
suspension feeding
what kind of fins do lamprey have
medial fins (paired fins and limbs absent)
when do the earliest vertebrates come from
early cambrian of China
what does the word conodont mean
cone teeth
what kind of feeding did conodonts have
selected and fed on larger food particles (prey)
when were ostracoderms most dominant
very late cambrian, silurian, early devonian
what was the first agnathan group to possess paired appendages
ostracoderms
what was the first vertebrate to have an intricate lateral line system
ostracoderms
what was the first vertebrate to have bone
ostracoderms
how large were most ostracoderms
minnow sized
what was the ostracoderm head shield
large bony plates of the head that were fused into a shield
what is the endoskeleton of ostracoderms made of
most fossils suggest cartilagenous endoskeletons
what is of interest in the fins of ostracoderms
paired muscular fins observed in the same position and vasculature as gnathostomes
what kind of falttening did ostracoderms have
dorsoventral
what are the three ostracoderm clades
pteraspidomorphs, osteostracans, anapsids
when do pteraspidomorphi ostracoderms appear
ordovician to late devonian
how many semicircular canals do pteraspidomorphi of the ostracoderms have
possess 2 semicircular canals
how many nares did pteraspidomorphi ostracoderms have
2
what did pteraspidomorphi never have and what did they sometimes have
sometimes had lateral and dorsal spines, and never has paired fins
ostracoderms body shapes were either fusiform or flattened, what does fusiform mean
spindle shaped
what was a popular tail morphology within the ostracoderms
hypocercal tail (bottom lobe longer than upper lobe)
what does the tail morphology of ostracoderms suggest
suggests ostracoderms did more open water swimming
what are the two early groups of jawed fishes
placoderms and acanthodii
when did acanthodii first appear
early sillurian
when did placoderms first appear
early sillurian
what was the early function of jaws
grasp, bite, crush prey (allowed for capturing of larger prey)
early gnathostomes has two types of paired fins, what were these fin types
pectoral and pelvic fins
where were the pectoral fins located on early gnathostomes
anterior
where were the pelvic fins located on early gnathostomes
posteriorly
what did paired fins allow for
stability and control
gnathostomes produced two major extant vertebrate groups, what are these two groups
chondrictes, teleostomi
what is within the chondrictes
sharks, skates, rays
what is within the teleostomi
bony fishes
when do placoderms date from
early silurian, flourished in devonian, gone in early carboniferous
what is different about the bone in ostracoderms versus placoderms
placoderms dont have dentin in their dermal bone
what distinguishes placoderms as a clade
bony thorax articulated with a bony head shield
what was the first fish to have internal bone
placoderms had the first internal bone (neural and hemal arches were ossified)
what was the size range for placoderms
hand sized to 30 feet long
what was probably the lifestyle of placoderms
benthic bottom feeders marine and freshwater
what are pelvic claspers
pelvic fins associated with internal fertilization (essentially penises)
placoderms were vivaparous, what does vivaparous mean
internal fertilization and give birth to live young
what group has the distinction of being the only major group of vertebrates to become entirely extinct without descendents
placoderms
what are within the elasmobranchs
(of chondrictes) its the sharks and the rays
what is within the holocephalans of chondrictes
chimearas (ratfish)
what is notable about the teeth of members in chondrictes
the teeth have serial replacement
what kind of scales do chondrictes have
placoid
when did chondrictes arise
early devonian
what kind of tail did placoderms have
heterocercal
why is it important to note the tail type of placoderms
heterocercal tail meant probably didnt have a swim bladder
what arch becomes the upper jaw
epibranchial cartilage of the first arch becomes the palatoquadrate cartilage of the upper jaw
what arch becomes the lower jaw
ceratobranchial cartilage of the first arch became mandibular cartilage (mekels)
what was the purpose of the second arch in the formation of jaws
in mammals it became hyoid arches, in other vertebrates it holds the first arch in place
if an organism retains the rest of its arches, what does it use them for
uses them to keep the cavity open and allows for gills
sharks have a spiracle on their head, what is this remenant of
the second pharyngeal opening moving upward creating a non functioning spiracle
chondrictes have sleek fusiform bodies, what does this suggest for function
suggests that they are/were active swimmers
what helps to keep chondrictes afloat
heterocercal tail and large liver filled with bouyant oils
what kind of birthing do chondrictes have
some have live birth, some have eggs
what is the functional support of most chondrictes
cartilagenous vertebrae
how many pharyngeal arches are in verts
7
how many pharyngeal openings are in verts
8
what group do rays belong to
batoidea
when does the ray fossils begin
early jurassic
what is the function of a spiracle in rays
primary way rays bring water into the body
what is the difference between skates and rays
skates have a rostum, rays dont