Exam 2 Flashcards
what is osteogenesis
the formation of bone
what are the two types of osteogenesis
intramembranous
cartilage replacement
what is osteogenesis in the form of intramembranous bone formation
happens to flat bone of skull and clavicles, happens within fibrous connective membrane
what is osteogenesis in the form of cartilage replacement bone formation
happens in most skeletal elements, can be perichondral or endochondral
what is an ossification center
the site of bone formation
what engages osteogenesis
osteoblasts
what removes existing bone
osteoclasts
what maintains fully formed bone
osteocytes
in what kind of development is the first formative bone nonlamellar
endochondral and intramembranous
what is another name for nonlamellar bone
immature bone or woven bone
what is nonlamellar bone
lots of cells interspersed amongst irregularly strewn bundles of collagen
what is perichondral
on the outside
what is endochondral
on the innermost
what is lamellar bone
mature bone
what does osteogenesis start with
local aggregations of loosel arranged mesenchymal cells
in what kind of osteogenesis formation is cartilage an intermediate
endochondral
when someone stops growing what has happened
their epiphysial plates have closed
what are the three regions have a bone
epiphysis, metaphysis, diaphysis
what is another name for the metaphysis
epiphyseal plate
what is the first step of bone formation from endochronral
loose cellections of mesenchymal cells consense to form hyaline cartilage
what is the second step of bone formation in endochondral
bone collar forms in region of diaphysis
what deposits the bone collar
cells on inner surface of diaphysis become osteoblasts and deposit the collar
what is happening in the bone at the same time as the formation of the bone collar
inorganic calcium salts accumulate in the matrix to calcify cartilage in the core of the diaphysis
what is the function of calcium in bone formation
calcify cartilage, seal off chondrocytes
what is the function of sealing off chrondrocytes in bone formation
seal them off so they die so vasculature can invade
when is the primary center of ossification established
when osteoblasts appear in the core of the bone
what happens in the center of ossification
old bits of calcified cartilage bone become overlaid by new bone
what are trabeculae
transitional composites of new bone
when are trabeculae considered bone spiracles
when an ossified matrix predominates
what do osteoclasts signal in osteogenesis
signals active nature of bone remodeling through matrix deposition and removal
what is the difference between osteoblast and osteoclast
blast deposits, clast removes
where is the active area of cartilage growth, calcification, cartilage removal, and new bone deposition
epiphyseal plates
what do chondrocytes do in response to approaching ossification
chondrocytes proliferate and hypertrophy as surrounding matrix calcifies
what lengthens bone
proliferation of cartilage in the epiphyses
what contributes to an increase in bone girth
continued depositon of bone under diaphyseal periosteum
why can fish grow larger even if reached sexual maturity
only bird and mammals stop growing when they reach adult and sexual maturity
what is the basic steps of osteogenesis
cartilage calcifies, blood vessels invade epiphyses, osteoblasts appear, new bone deposited
when do secondary centers of ossification appear in humans
at 2-3 years of age
when does the zone of ossification take over cartilage proliferation
at or shortly after mammals reach sexual maturity
what are the 5 chordate characteristics that all vertebrates have
notochord, pharyngeal openings, post anal tail, endostyle, dorsal hollow nerve cord
what are the innovations of chordates that are characteristic to vertebrates
cranium and vertebrae
what are vertebrae
series of separate bones or cartilage blocks firmly joined together as a back bone that defines the major axis of the body
what are between vertebrae
intervertebral disks
what are the 3 basic parts to a vertebra
centrum, neural arch, hemal arch
what structural element did early vertebrates rely on
has vertebrae but still mainly relied on the notochord
where does the notochord remain in advanced vertebrates
in the nucleous pulpous in the intervertebral disks
what is the function of a cranium
supports sensory organs in the head and encases or partially encases the brain
what does cephalization mean
anterior clustering of specialized sensory organs
what are the three parts of the brain
forebrain, midbrain, hindbrain
what two kinds of cells give rise to most sensory organs in the head and are the source of most adult structures that distinuish vertebrates from chordates
neural crest cells, epidermal placodes
in what habitat are the earliest vertebrates found
marine waters
what are the three stages of vertebrate evolution
prevertebrates, agnathans, gnathostomes
what is a prevertebrate
suspension feeding organism that probably resembled amphioxious without burrowing habits
how did agnathans feed
use of a muscular pump to produce food bearing water current
where did the prevertebrates arise from
arose within protochordates
what two mechanical changes caused the shift from prevertebrate to vertebrate
pharynx developed encircling band of muscles, strong and springy cartilage replaced collagen in pharyngeal bars (this formed the muscular pump)
what removed the limits to size in early vertebrates
the loss of cilliary pump and the dominance of muscular pump
what did the muscular pump of prevertebrates allow for
allowed for larger body size, more complex feeding, addition of gills instead of pharyngeal openings
what are gills
complex, folded respiratory organs on pharyngeal pouches whose folds contain blood capillary beds capable of gas exchange
what are the folds of gills called
lamellae
what did the addition of gills allow for in vertebrates
allowed for increased respiratory demands made by organism due to increased activity
how did early vertebrates arise to the agnathan stage
appearance of muscular pharyngeal pump
what are the 5 agnathan groups
conodonts, pteraspidomorphi, ostracoderms, myxini, pteromyzontida
what kind of feeding did early gnathostomes posses
raptiorial and suction feeding
what removed size restriction of prey for gnathistomes
addition of jaws
if youre not a fish youre a ____
tetrapod
what are amniotes
vertebrates that wrap their embryos in an amnion
who are the anamniotes
fishes and amphibians
what are jaws derived from
phyrengeal arches
were ostracoderms vertebrates
no, notochord instead of vertebrae
ostracoderms were not considered craniates, why
they had a dermal bone head shield but not a chondral cranium
what do odontoblasts do
form dentin and enamel (mesenchymal derivatives)
how does dermal bone vary most significantly from other bone types
dermal bone has an enamel surface
what is the issue with having bone made of enamel and dentin
no blood vessels so its not reparable
what are the extant agnathans
hagfish and lampreys (myxini and pteryomyzontida)
what kind of larvae do lampreys have
ammocoete (kind of like amphioxious)
what kind of teeth do lampreys have
keratinized teeth (like finger nails)
lampreys and hagfish do not have arcualia when embryonic, what are arcualia
theyre embryonic structures that form vertebrae
whats the ecological difference between hagfish and lampreys
lampreys are ectoparasites
what kind of feeding do hagfish have
carrion feeding
what is carrion feeding
eat dead stuff
how many semicircular canals do lampreys have
2
how many semicircular canals do hagfish have
1
hagfish are unique in that they are isoosmotic, what is isoosmotic
organisms cells are in balance with the salt of the ocean
what organism displays knotting feeding when feeding
hagfish
what are the cyclostomes
hagfish and lamprey because they have circular mouths
where does the fossil data for hagfish and lampreys reach to
late devonian
all agnathans lack ____ and possess____
lack bone and possess a single nostril
what is the function of hagfish knotting
provides force to tear off food from a larger body
hagfish are hermaphrodites but they are not practicing hermaphrodites, what does this mean
they have both sets of gonads but only use one set, must have another hagfish present to reproduce
what kind of agnathan has vertebrae
some hagfish have embryonic vertebrae
what does it mean for water movement in the body of the hagfish that it is isoosmotic
no net flow o water in or out of the body (doesnt have to drink or pee)
what was most likely the direct ancestor to hagfish
invertebrates
how does a lamprey stay in place in moving water
grasps rock in mouth and holds it
what kind of feeding does the lamprey larva show
suspension feeding
what kind of fins do lamprey have
medial fins (paired fins and limbs absent)
when do the earliest vertebrates come from
early cambrian of China
what does the word conodont mean
cone teeth
what kind of feeding did conodonts have
selected and fed on larger food particles (prey)
when were ostracoderms most dominant
very late cambrian, silurian, early devonian
what was the first agnathan group to possess paired appendages
ostracoderms
what was the first vertebrate to have an intricate lateral line system
ostracoderms
what was the first vertebrate to have bone
ostracoderms
how large were most ostracoderms
minnow sized
what was the ostracoderm head shield
large bony plates of the head that were fused into a shield
what is the endoskeleton of ostracoderms made of
most fossils suggest cartilagenous endoskeletons
what is of interest in the fins of ostracoderms
paired muscular fins observed in the same position and vasculature as gnathostomes