Chapter 44 Flashcards
What are exteroceptors?
receptors that sense stimuli that arise in the external environment
Evolutionarily speaking, where did vertebrate sense evolve?
first in water before the invasion of land
What are the consequences of the fact that vertebrate sensory systems evolved in water before land?
may senses of terrestrial vertebrates emphasize stimuli that travel well in water
What are interoceptors?
receptors that sense stimuli that raise from within the body
Which are typically more complex, exteroceptors or interoceptors?
exteroceptors
Which bear more of a resemblance to primitive sensory receptors, exteroceptors or interoceptors?
interoceptors?
What are the three broad classes of receptors?
mechanoreceptors, chemoreceptors, electromagnetic receptors
What are mechanoreceptors? (2)
stimulated by mechanical forces such as pressure; include receptors for touch, hearing, and balance
What are chemoreceptors? (2)
detect chemicals or chemical changes; includes receptors for smell and taste
What are electromagnetic receptors? (2)
react to heat and light energy; includes eye photoreceptors
What is the four-step process for the conveying of sensory information?
stimulation, transduction, transmission, interpretation
Describe stimulation in the four-step process for the conveying of sensory information.
a physical stimulus impinges on a sensory neuron or associated/separate sensory receptor
Describe transduction in the four-step process for the conveying of sensory information.
stimulus energy is transformed into graded potentials in the dendrites of the sensory neuron
Describe transmission in the four-step process for the conveying of sensory information.
action potentials develop in the axon of the sensory neuron and are conducted to the CNS along an afferent nerve pathway
Describe interpretation in the four-step process for the conveying of sensory information.
brain creates sensory perception from the electrochemical events produced by afferent stimulation.
Why do sensory cells respond to stimuli?
because they possess stimulus-gated ion channels in their membranes
In most causes, the sensory stimulus causes what event in the receptor cell?
causes a depolarization f the receptor cell, analogous to an EPSP
What is a receptor potential?
a depolarization that occurs in a sensory receptor on stimulation
What type of potential is a receptor potential?
a graded potential
What is the relationship between the size of the sensory stimulus and the degree of polarization?
the larger the sensory stimulus, the greater the degree of polarization
What is responsible for conveying the intensity of a stimulus?
the frequency of action potentials, NOT their summation
What is the relationship between stimulus intensity and action potential frequency?
a logarithmic relationship
What is the name for the receptors of the skin?
cutaneous receptor
Cutaneous receptors are classified as
interoceptors
What are nociceptors?
the receptors that transmit impulses perceived as pain
Most nociceptors consist of
free nerve endings located throughout the body, especially near surfaces where damage is most likely to occur
What is transient receptor potential (TRP) ion channel?
a class of ion channel proteins found in nociceptors
The TRP ion channel is stimulated by
temperature to produce an inward flow of cations of Na+ and Ca2+
The inward flow of Na+ and Ca2+ through the TRP ion channel causes
a depolarizing current which further causes the sensor neuron to fire, leading to the release of glutamate and an EPSP in neurons in the spinal cord, ultimately producing the pain response.
The first identified TRP channel responds to
the chemical capsaicin, which is found in chili peppers, and also responds to heat
A cold-responsive TRP receptor responds to
the chemical menthol
How do TRP channels reduce the body’s pain response?
chemical stimulation of the TRP channels reduces the body’s pain response by desensitizing the sensory neurons
What are thermoreceptors?
naked dendritic endings of sensory neurons that are sensitive to changes in temperature
How are nociceptors and thermoreceptors similar?
nociceptors also contain free nerve endings
Thermoreceptors contain
TRP ion channels that are responsive to both hot and cold
Where are cold receptors located?
immediately below the epidermis
Which are more numerous, cold or warm receptors?
cold receptors are 3-4 x more numerous than warm receptors
Where are warm receptors located?
slightly deeper in the dermis, below the cold receptors
Where are thermoreceptors found in the brain?
in the hypothalamus, where they monitor the temperature of circulating blood and provide the CNS with info on the body’s core temp
What are four main types of mechanoreceptors in the skin?
merkle cell, meissner corpuscle, ruffini corpuscle, pacinian corpuscle
What are Merkle cells?
tonic receptors located near the surface of the skin that are sensitive to touch pressure and duration
What are Meissner corpuscles?
phasic receptors sensitive to fine touch, concentrated in hairless skin
What are Ruffini corpuscles?
tonic receptors located near the surface of the skin that are sensitive to touch pressure and duration
What are Pacinian corpuscles?
pressure-sensitive phasic receptors deep below the skin and in the subcutaneous tissue
What are the two tonic receptors?
Merkle cells and Ruffini corpuscles
What are the three phasic receptors?
Meissner corpuscles, hair follicle receptors, and Pacinian corpuscles
The four mechanoreceptors in the skin contain
sensory cells with ion channels that open in response to mechanical distortion of the membrane
What does it mean to be phasic?
intermittently activated
What does it mean to be tonic?
continuously activated
Phasic receptors are found on surfaces
that do not contain hair, like fingers/palms/nipples
Pacinian corpuscles consist of
the end of an afferent axon surrounded by a capsule of alternating layers of connective tissue cells and extracellular fluid
What happens when sustained pressure is applied to a Pacinian corpuscle?
the elastic capsule absorbs much of the pressure and axon ceases to produce impulses
Muscle length and tension are monitored by
proprioceptors
What are muscle spindles?
sensory stretch receptors that lie in parallel with the rest of the fibers in the muscle
Each muscle spliced consists of
several thin muscle fibers wrapped together and innervated by a sensory neuron, which becomes activated when the muscle/spindle is stretched
What are proprioceptors?
sensory receptors that provide information about the relative position or movement of the animal’s body parts
What are Golgi tendon organs?
a proprioceptor that monitors the tension exerted on tendons when an associated muscle contracts
What do baroreceptors do?
detect blood pressure
Blood pressure is monitored at what two main sites in the body?
the carotid sinus and the aortic arch
What is the carotid sinus?
an enlargement of the left and right internal carotid arteries that supply blood to the brain
What is the aortic arch?
the portion of the aorta very close to its emergence from the heart
What exactly do baroreceptors detect?
they detect tension or a stretch in the walls
Describe what happens when blood pressure decreases. (2)
the frequency of impulses produced by the baroreceptors decreases; CNS responds by stimulating sympathetic division which increases heart rate and causes vasoconstriction
Describe what happens when blood pressure increases. (2)
baroreceptor impulses increase; reduces sympathetic activity and instead stimulates parasympathetic division which lowers heart rate
What is hearing?
the detection of sound waves
Why does hearing work better in water than in air?
because water transmits pressure waves more efficiently
What system in fish provides a sense of “distant touch?”
lateral line system
The lateral line system enables fish to
sense objects that reflect pressure waves and low-frequency vibrations
Is the lateral line system found in amphibians?
found in amphibian larvae but is lost at metamorphosis
Is the lateral line system found in terrestrial vertebrates?
no
How does the lateral line system interact with the fish’s hearing?
the lateral line system supplements the fish’s sense of hearing
What does the lateral line system consist of?
hair cells within a longitudinal canal in the fish’s skin that extends along each side of the body and within several canals in the head
(lateral line system) The hair cells’ surface processes project into
a gelatinous membrane called a cupula
(lateral line system) hair cells have hairlike processes called
stereocilia
What is a kinocilium?
longer hairlike process present on hair cells in the lateral line system
Describe sterecilia. (2)
actually microvilli; contain actin fibers
Describe a kinocilium. (2)
true cilium; contains microtubules
If the stereocilia are bent in the direction of the kinocilium, what happens?
associated sensory neurons are stimulated and generate a receptor potential
If the stereocilia are bent in the opposite direction of the kinocilium, what happens?
activity of the sensory neuron is inhibited
Otoliths are composed of
calcium carbonate crystals
Where are otoliths located?
otolith organs of the membranous labyrinth, a system of fluid-filled chambers and tubes also present in other vertebrates
What structure in the fish is primarily responsible for detecting sound?
otoliths
What structures make up the outer ear? (2)
ear canal; ear drum (tympanic membrane)
Vibrations of the tympanic membrane cause movement of
one or more small bones that are located in a bony cavity known as the middle ear
Amphibians/reptiles have what single middle ear bone?
stapes (stirrup)