Topic 3: Exchange + Transport: Transport in Plants Flashcards
Why do multicellular plants need transport systems?
Multicellular plants need transport systems because:
-Metabolic demands = underground parts of plants don’t photosynthesise but still require exchange of substances and transportation of hormones and mineral ions absorbed by roots
-Size = plants grow large -> needs effective transport systems to move substances up and down between tips and roots
-Surface area = small sa:vol ratio -> cannot rely on diffusion alone to supply cells
What are dicotyledonous plants (dicots)?
Dicotyledonous plants make seeds that contain two cotyledons (organs that act as food stores for the developing embryo plant and form the first leaves when the seed germinates)
What are cotyledons?
Cotyledons are organs that act as food stores for the developing embryo plant and form the first leaves when the seed germinates
Types of dicots (dicotyledonous plants)
Types of dicots:
-Herbaceous dicots
-Woody (arborescent) dicots
Herbaceous dicots
Herbaceous dicots - type of dicot - soft tissues and relatively short life cycle (leaves and stems that die down at the end of the growing season to the soil level)
Woody (arborescent) dicots
Woody (arborescent) dicots - type of dicot - hard, lignified tissues and a long life cycle (sometimes hundreds of years)
Vascular systems in dicotyledonous plants
-Vascular system: dicots have a series of transport vessels running through the stem, roots and leaves which is known as the vascular system
Vascular bundles in dicotyledonous plants
-Vascular bundles: transport tissues (eg phloem + xylem) are arranged together in vascular bundles in leaves, stems and roots of harbaceous dicots
Why are vascular bundles around the edge in stems?
Vascular bundles are arranged around the edges in stems to give strength and support
Why are vascular bundles in the middle of roots?
Vascular bundles are in the middle of roots to help the plant withstand tugging strains that result as the stems and leaves are blown in the wind
The midrib of a dicot leaf
The midrib of a dicot leaf is the main vein carrying the vascular tissue through the organ. It also helps to support to the structure of the leaf. Many small, branching veins spread through the leaf functioning both in transport and support.
Function of the xylem
Function of the xylem: large non-living tissue that has two main functions - the transport of water and mineral ions, and support
-The flow of materials in the xylem is up from roots to shoots and leaves
-Made up of dead cells
-Long, hollow structures made by columns of cells fusing together end to end
Two other tissues associated with the xylem in herbaceous dicots
Two other tissues associated with the xylem in herbaceous dicots:
-Thick-walled xylem parenchyma packs around the xylem vessels, storing food and containing tannin deposits
-Xylem fibres with lignified secondary walls that provide extra mechanical strength but doesn’t transport water
Tannin in dicots
Tannin is a bitter, astringent-tasting chemical that protects plant tissues from attack by herbivores
Xylem parenchyma in dicots
Thick-walled xylem parenchyma packs around the xylem vessels, storing food and containing tannin deposits
Xylem fibres in dicots
Xylem fibres in dicots:
Long cells with lignified secondary walls that provide extra mechanical strength but do not transport water
Ways of which lignin can be laid down in the walls of xylem vessels
Ways of which lignin can be laid down in the walls of xylem vessels:
Rings, spirals or relatively solid tubes with lots of small unlignified areas called bordered pits
Bordered pits in plants
Bordered pits is where water leaves the xylem and moves into other cells of the plant
Functions of the ploem
-Phloem is living tissue that transport food in the form of organic solutes around the plant from leaves where they are made by photosynthesis
-Supplies cells with sugars and amino acids needed for cellular respiration and for synthesis of all other useful molecules
-Flow of materials in phloem can go both up and down the plant
Sieve tube elements in phloems
Sieve tube elements - the main transporting vessels of the phloem - made up of cells joined end to end forming a hollow structure
-Not lignified
Sieve plates in the phloem
Sieve plates - areas between cells where walls have become perforated (hole-y) - tonoplants (vacuole membrane), nucleus and some organelles break down - phloem becomes tube filled with sap and mature phloem cells have no nucleus
Companion cells in the phloem
Companion cells form along with sieve tube elements - cells are linked to the sieve tube elements by many plasmodesmata - maintains nucleus and organelles
-Active cells and function as ‘life support system’ for sieve tube cells, which have lost normal cell functions
What is plasmodesmata?
Plasmodesmata are microscopic channels through the cellulose cell walls linking the cytoplasm of adjacent cells
Tissues that support the phloem tissue
Fibres and scelerids (cells with with extremely thick cell walls) are supporting tissues found within the phloem
What are sclereids?
Scelereids are cells with extremely thick cell walls, and act as supporting tissue (eg they are found in the phloem)
Vascular cambium in stems
Vascular cambium in stems - located inbetween the xylem and phloem cells - contains meristem cells
Arrangement of xylem and phloem in stems
Stem arrangement: circular, xylem and phloem vessels located around it, with xylem being inside and phloem being on the outside, vascular cambium located inbetween the xylem and phloem vessels and contain the meristem cells
Arrangement of xylem and phloem in roots
Xylem is located at the center of roots in an ‘X’ drill-like structure, with phloem in four seperate sections around the xylem
Why is water important in both the structure and in the metabolism of plants?
Water is important in both the structure and metabolism of plants because:
-Turgor pressure (or hydrostatic pressure) as result of osmosis in plant cells provides a hydrostatic skeleton to support the stems and leaves (around 1.5MPa in plants)
-Turgor drives cell expansion - force enables plant roots to force their way through tarmac and concrete
-Loss of water by evaporation helps to keep plants cool
-Mineral ions and the products of photosynthesis are transported in aqueous solutions
-Water is a raw material for photosynthesis
Root hair cells function
Root hair cells are the exchange surface in plants where water is taken into the body from the plant from the soil
-Root hair is long, thin extension from root hair cell
How are root hairs well adapted as exchange surfaces?
Root hair adaptations:
-Microscopic size = penetrate easily between soil particles
-Each microscopic hair = large sa:v ratio
-Each hair has thin surface layer which means diffusion and osmosis can take place quickly
-Concentration of solutes in the cytoplasm of root hair cells maintains a water potential gradient between the soil water and the cell
Why does soil water have a high water potential?
Soil water has a high water potential due to having a low concentration of dissolved minerals
Why does the cytoplasm and vacuolar sap of the root hair cell have low water potential?
The cytoplasm and vacuolar sap of the root hair cell has a low water potential due to containing many different solvents including sugars, mineral ions and amino acids - as a result water moves into the root hair cells by osmosis
Three different pathways that water can move across the root to the xylem once it has entered the root hair cell?
Two pathways water can go leading up to the xylem vessel:
-The symplast pathway
-The apoplast pathway
-Vacuolar pathway
What is the symplast of plants?
The symplast is the continuous column of living plant cells that is connected through the plasmodesmata
What is the apoplast in plants?
Apoplast in plants: the cell walls and the intercellular spaces of plants
Movement of water into the xylem
Movement of water into the xylem:
Water moves across root (apoplast or symplast pathway) until epidermis -> water in the apoplast pathway forced into cytoplasm and joins water in symplast pathway, passing through a selectively permeable membrane -> once inside vascular bundle, water returns to apoplast ptathway to enter xylem
Why does water in the apoplast pathway pass through selectively permeable membranes before entering the symplast pathway on the route to xylem vessels?
passing through selectively permeable cell surface membranes excludes potentially-toxic solutes in soil water from reaching living tissues
What causes root pressure?
The active pumping of minerals into the xylem to produce movement of water by osmosis results in root pressure - independant of any effects of transpiration
Root pressure
Root pressure gives water a push up the xylem, but under most circumstances it is not the major factor in movement of water up from roots to leaves
Evidence for the role of active transport in root pressure
Evidence for the role of active transport in root pressure:
-Some poisons, eg cyanide, affect the mitochondria and prevent ATP production - if cyanide is applied to root hair cells there is no energy supply and root pressure disapears
-Root pressure increases with a rise in temp and falls with a decrease in temp, suggesting enzyme controlled reactions are involved
-Levels of oxygen or respiratory substrates fall - root pressure falls
-Guttation: sap and water moves out cut stems, suggesting they are actively pumped out rather than being drawn out by transpiration