Seeds Flashcards
What is the definition of a seed?
A seed is a reproductive structure (propagule) which is formed by the maturation of the ovule of seed plants, following fertilization
What are the advantages of seeds?
- Seed coat (testa) provides protection e.g. from mechanical damage.
- More reserves than a spore - endosperm or cotyledon provides nourishment for the seedling while it germinates.
- Seeds can lie dormant until conditions for growth are more favourable (survive - winter, drought)
- Seeds usually provide a repository of genetic variation (meiosis, cross pollination)
- No genetic variation in cuttings and tuber - Means of dispersal – animals, wind, projectile, water..
What makes up a seed?
• seed = embryo + seedcoat (testa) OR seed = embryo + testa + endosperm
• embryo = plumule + radicle + cotyledon(s)
• plumule : the first bud of an embryo; part of the
embryonic axis above the cotyledonary node, almost the
same as epicotyl
• radicle : embryonic root
• hypocotyl: region below the cotyledon node and true root
is where stem & root anatomy „swap over‟
• cotyledon :first leaf of an embryo; 1 (monocot) or 2
(dicot) per seed; functions in:
– foodstorageor
– absorptionfromtheendospermor
– photosynthesis
What are the comparisons between monocot and dicot seeds?
“Grasses”
• Onecotyledon • • Starch stored outside the •
embryo (endosperm) • Hypogealgermination
“Broad leafs” Twocotyledons
Energy stored within the embryo (most cases)
• Bothhypogealand epigeal germination
What conditions are needed for seed germination?
- Adequate water
- Suitable temperature
- Adequate oxygen
- Adequate light level (if needed)
- Absence of inhibitors/toxins
Adequate Water
Rate at which seeds take up water (imbibition) based upon:
1.Permeability of the seed coat (testa)
scarification (scratching of water impermeable seed coat – e.g. your Acacia seeds)
low temperature treatment (stratification)
2.Seed composition.
starchy seeds take up water quickly (corn or wheat) oily or fatty seeds take up slower (canola or peanut)
3.Whether the water is in the form of: liquid
vapour (high humidity)
Suitable Temperature
Based upon 3 “CARDINAL” temperatures
1. Minimum temperature below which no germination
2. Optimal temperature = highest % germination in the shortest time
3. Maximum temperature above which no germination
Beyond the maximum are lethal temperatures = seed death (~50oC)
Oxygen
- C6H12O6 +O2 → CO2 + H2O + “ENERGY” • respiration „reverse‟ of photosynthesis
- aerobic vs. anaerobic respiration
- rate of respiration dependent upon the water content of the seed
Light
• Most seeds are light independent relative to germination
• Exceptions:
– germinate only in the dark
– germinate only in the light
– germinate only after a short light exposure
– germinate only after illumination with Far-Red light
Light and Temperature Interactions
• Light or temperature alone may be dangerous as germination triggers
• Combinations often allow for more ideal growth conditions
• Small seeds (e.g. Canola)
– Often sown shallow because of:
• Small energy reserves (barely enough energy to reach the light)
• Requirement for light to germinate • Which came first?
Inhibiting Conditions
• Internal
– Compounds present in the seed coat
• External
– Gases – excess CO2 – Salts
– Chemicals
Most of these can be washed or LEACHED away by large amounts of water.
• Water, temperature, O2 and no toxins/inhibitors seeds still may not germinate
• termed dormancy
– Imposed by the seed coat (physical and/or physiological
– Imposed by the embryo (physiological)
What does Seed Coat Dormancy mean?
• Impermeable to Water (physical) • Impermeable to Oxygen and CO2 (physical) • Mechanical Resistance (physical) • Germination-inhibiting chemicals (physiological)
How is Seed Coat Dormancy overcome?
• Human intervention
– Cutting / scratching the seed coat – Chemical (acid) treatment
• Natural means
– Scarification (scratching) of the seed coat by soil or animals etc.
– Freeze-thaw cycles
– Fire
– Cycles of water to leach out inhibitors
What does Physiological Dormancy mean?
- Embryo is not yet fully developed (e.g. orchid seeds require symbiotic fungi)
- Requirement for energy storage
- Requirement for light (specific wavelength)
- Requirement for chilling (stratification) (Hormones: Abscisic acid deactivation, Gibberellin synthesis)
- Combinations of some or all of the above
What are some advantages of seed dormancy?
- Can delay germination until environmental conditions are favourable.
- Creation of a seed bank
- Can help synchronize germination to a particular time of the year
- Can facilitate seed dispersal