Primary and Secondary Growth Flashcards
root primary growth
root apical meristem (RAM)
- composed of apical meristem in centre of root tip
subdivided into 3 meristematic areas:
- protoderm
- ground meristem
- procambium
protoderm
gives rise to epidermis
ground meristem
gives rise to cortex (parenchyma, exodermis, endodermis)
procambium
gives rise to primary xylem, primary phloem, and pericycle (vascular cylinder)
region of maturation
cells mature (differentiate) into distinctive cell types
region of elongation
cells become several times original length; not differentiated (yet)
pericycle
single cell layer, located directly below endodermis
- gives rise to lateral roots and part of vascular cambium
lateral root development: pericycle
- pericycle displaces cortex and epidermis as lateral root grows outward, from vascular cylinder to epidermis
- lateral root eventually breaks through epidermis
- vascular cylinder (xylem and phloem) of lateral root stays connected to that of parent root
secondary root growth
- no secondary root growth in fibrous monocot root systems
- as in stems, secondary growth occurs where primary growth has been completed
- dicot and conifer taproots have secondary growth to:
-increase conductivity
-replace old vascular tissue
-provide protection (periderm=bark)
-form specialized storage roots by anomalous secondary growth
-provide support in specialized roots
2 types of lateral meristems
present in both dicot and conifer stems and roots
vascular cambium and cork cambium
secondary root growth: vascular cambium
- forms from dedifferentiated parenchyma between primary xylem and phloem and pericycle
- initiated by auxin
- forms continuous ring of meristematic cells
as in stems, secondary xylem = wood and less secondary phloem is produced
secondary root growth in dicot and conifer: cork cambium
- forms from dedifferentiated parenchyma cortex, then secondary phloem
secondary root growth in dicto and conifer: annual growth rings
roots also have growth rings but harder to distinguish as secondary xylem sizes do not vary by growing seasons
anomalous secondary thickening: food storage
- storage cells of root vegetables like beets, carrots, radishes are a combination of root and stem, and store simple carbohydrates
- storage root expands by anomalous secondary thickening of xylem, phloem, and parenchyma
- beets have growth rings, unlike in stem where only secondary xylem, rings are comprised of repeating secondary xylem, secondary phloem, and storage parenchyma
root periderm scars
- many food storage tap root shave root periderms scares resembling lenticels but NOT for gas exchange
- when lateral root breaks through periderm, wound response creates scarring