Lecture Week 3 Flashcards
Auditory Sense of Self
- Sense of Sound contributes to our sense of self
- Playing sounds of a certain heart rate; if a person thinks that is their own heart rate then their own heart rate will change to match the recording.
Physical Definition of Sound
- Pressure changes in the air or other medium
- Could also be pressure change in water, a gas or a solid
Perceptual Definition of sound
- The experience we have when we hear.
How is sound created?
- Sounds are created when objects vibrate
- Vibrations cause the surrounding air to change pressure
- The type of pressure change determines the type of sound wave
- The sound wave determines what we hear
Amplitude Waves
- The Level of loudness of a sound
- Objects that vibrate a lot will be perceived as loud
- Measured in Decibels dB
Frequency Waves
- The number of times per second that a sound wave repeats the pattern of pressure
- The number of times the waves go up and down per second
- Measured as Hz - Hertz
- Perceived as pitch; High frequency is high pitch and low frequency is low pitch
High Risk and Pain Threshold in Human Hearing
- At around 115 dBs we begin to experience sound that may cause us damage.
- At around 135 dBs our nociceptors will begin to transduce and we will experience pain.
- This is about equivalent to the sound of a gun going off next to your ear.
Hyperacusis
The hearing condition with lingering pain
Psychological Interpretation of Sound - Loudness
- How we experience sound of a particular amplitude
- High amplitude is loud
- Low amplitude is quiet
Psychological Interpretation of Sound - Pitch
- Our experience of sound related as a frequency
- High frequency, High pitch, High Hertz
- Low frequency, Low pitch, Low Hertz
Psychological Interpretation of Sound - Timbre
- The ability to judge two sounds with the same amplitude and frequency as being different from one another
- Like the flavour of the music and not the taste
eg: the same note being played by a piano and a guitar
Outer Ear
- Also called Pinnae are first point of contact of sound from the environment
- Only Mammals have outer ears
- Some mammals do not have pinnae such as seals and Walruses
- They funnel sound into the ear canal
Ear Canal
- Insulates and protects the tympanic membrane
- Also funnels sound and serves to amplify some frequencies
Tympanic Membrane
- Also called the Ear Drum
- A barrier between outer ear and middle ear
- It is made of thin skin so if it is damaged it will repair like any other peice skin
- Vibrates in response to fluctuations in air pressure
- When it vibrates it moves the bones in the inner ear
Middle Ear
- Consists of three very small bone called ossicles
- Malleus
- Incus
- Stapes
- Two small muscles
- Tympanus Muscle
- Stapedius Muscle
Ossicles
- Purpose of Ossicles is to amplify sounds
- Smallest Bones in the Human Body
- Malleus
- Incus
- Stapes
- Ossicle amplicifaction is essential for hearing faint sounds they amplify sound x18
Malleus Ossicle
- Is connected to the Tympanic Membrane
- It moves when the Tympanic Membrane moves
- They move each other around in response to flucutations in air pressure
- They operate like levers
- Ossicle amplicifaction is essential for hearing faint sounds they amplify sound x18
Tensor Tympanic Muscle & Stapedius muscle
- Decrease Ossicle vibration when tensed
- They protect our ears from loud noises
- When they tense they hold the ossicles stable so they dont amplify sound
- Fast responding by 1/5 of a second
Inner Ear
- Transduces responses to air pressure changes into neural signals to be processed as sounds
Cochlea
- Contains the vestibular organs
- Organ of Corti
- semi circular canals
- Otolithic Organs
- Filled with watery cochlear fluid in all the canals
- When the ossicles vibrate this causes the cochlear fluid to vibrate as well
- This movement affects the:
- Basilar Membrane
- Tectorial Membrane
Organ of Corti
- Responsivbe for transducing changes in air pressure into neural activity
Basilar Membrane
- Where the Stereocilia are located
- This is specifically where the hair cells begin the process of transduction
- Hair cells respond to the movement in the cochlear fluid
Tectorial Membrane
- A gelatinous structure that extends into the midde canal of the ear
- It causes the hair cells to fluctuate and it depends on how the hair cells move on what type of signals get transduced
- This movement is called Tectorial Shearing.
Pitch and the Basilar Membrane
- Different parts of the cochlear are sensitive to different types of ptch
- This is because the cochlear is wider at one end than the other
- This changes the way the hair cells can bend or change the amount of stimulation they need to bend
Cochlear Nucleus
- The first brain stems nucleus at which afferent auditory nerve fibres synapse
- Then sends neural activity to the Superior Olive
Superior Olive
- Region of the brain stem where auditory input is received
- The first place where inputs from both ears are received
- Intergrates information so we can quickly locate where a sound is coming from
Inferior Colliculus
- Also important for Binaural Integration
- In the midbrain nucleus
- Then sends information to the Medial Geniculate Nuclesu of the Thalamus
Binaural Integration
Integrate and process neural information from both ears
Medial Geniculate Nuclesu of the Thalamus
- Thalamus is the relay station for all of our senses
- MGN sends auditory signals to the temporal cortex
- MGN receives signals from auditory cortex
- Information from both ears are processed here on both sides
Primary Auditory Cortex (A1)
- In the temporal lobe
- Sends information on to the belt area and parabelt area
- Responsible for acoustic organisation
Belt Area
- A region of the A1 Cortex
- Directly adjacent to the A1
- where neurons respond to more complex characteristics of sounds
Parabelt Area
- Lateral and Adjacent to the Belt Area
- Where neurons respons to more complex sounds as well as inputs from other senses
Left-Right Localisation
- Interaural Time Differences
- Interaural Level Differences
Azimuth
- The angle of a sound source on the horizon relative to a point in the centre of the head
- Anything that happens 360O on the Horizon is called Azimuth
- To work out where things are on the Azimuth we can use our ITDs and ILD’s
- We can also disambiguate using the shape of our Pinnae
Disambiguate in Hearing
- Remove uncertainty about sound using both ears to detect differences on the Azimuth
Interaural Time Differences
- The difference in time between a sound arriving in one ear over the other depends on which ear the sound is closest to.
- If a sound is straight ahead or straight behind the ITD will be 0
- If it is directly at either side it will be maximum ITD
- It is slightly easier to hear sounds from the front because the shape of our Pinnae
Medial Superior Olive
- Interaural Time Differences are processed here
- Happens from the first synapse to the Superior Olive to Thalamus
- Neurons from the left and right Olive work together to calculate time differences
- This assists in sound localisation
Interaural Level Difference
- Frequency levels will be different in one ear over the other.
- The Medial Superior Olive receives info from both ears and uses the differences to assist in localisation
- There will be different levels of frequency depending on how much the frequencies are blocked by our heads
- This works well for high frequencies
- Low frequencies are better at working around the head anyway.
- Excitatory Neural synapses from here lead to the Lateral Superior Olive from the ipsilateral ear.
- Inhibitiry Neural synapses from here lead to the Lateral Superior Olive from the contralateral ear.
Lateral Superior Olive
- A relay station that Calculates the frequency differences and ILDs from both ears to assist localisation
- Excitatory synapses come from the ipsalateral ear
- Inhibitory synapses come from the contralateral ear
Cone of confusion
- There are positions in space around the azimuth where the ITDs & ILDs are the same.
- In this case the Olives will tell us the location of two positions front and back are the same.
- These positions are situated at the same place at 180o in front and behind us.
Marco Polo Game as an example; these are the best places to stand and not get caught.
- Elevation adds another dimension to help distinguish this problem
- Turning our heads to disambiguate can help too.
Directional Transfer Function (DTF)
- How the shape of our pinnae, ear canal, head and torso can affect the intensity of sounds and frequencies arriving at each ear
- each person has a unique DTF to determine localisation based on the azimuth and elevation of that sound
Ventriloquism Effect
- Sound localisation can be influenced by visual stimulus.
- If the audio-visual stimulus is strong enough we can localise a sound as if it is coming from the visual space
eg: a ventriloquist dummy. The illusion still works even when we know it is happening,
Auditory Distance Perception
- The simplest cue is relative intensity of sound
- Sounds that are closer to us are louder than sounds that are far away.
- Works best when we are familiar with a sound and we know how loud or soft it should be
Relative Intensity of Sound
- Sounds that are closer to us are louder than sounds that are far away.
Inverse Square Law
- Relativity of Sound only works at distances of about 1 metre
- Decrease in intensity of a sound is equal to the Distance2 from the sound
- As an object moves away from us, its sound decreases faster.
- For this reason we can over and understimate the intensity of sounds as they move.
Auditory Scene Analysis
- The method to sort out multiple sources of sound in an environment
- Uses information from:
- visual stimulus
- Interaural Time Differences
- Interaural Level Differences
- Pitch
- Timbre
- Onset Time
Onset Time
- If two sounds start at different times it is likely they come from different sources
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Continuity and Restoration Effect
- We can have auditory restoration and phonemic restoration
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Restoration Effect
- Masking sounds and brain fills in gaps in sound so that it sounds continuous
- There has to be a masking sound other=wise the brain will not fill in the continuous sound
Articulation
- Converting the sounds made by the vocal tract into phonemes and speech
Formant
- Different parts of our articulation repeat at different frequencies
- This creates formants in the speech spectrum
Formants are listed by Number
- F1 tells about the height of the tongue in the mouth
- F2 tells about how forward or back the tongue is
- F3 is a combination of the above
- We can distinguish most sounds in the English language based on these three formants
Phonemes
- The sounds that make up words
- Almost like the letters but combinations such as th and ph are also phonemes
- The smallest unit of sound that has meaning
Classifying Speech sounds – Place of articulation
Based on the location in which the sound is made such as b, p, m are all lip sounds
Classifying Speech sounds – Manner of articulation
Depends on the manner and amount of air being obstructed to construct that sound
Categorical Perception
- Situation in which a speech stimulus is perceived in terms of its category membership
- Not in terms of its surface properties.
- We perceive an utterance as a specific syllable, word, or phrase.
Motor Theory of Speech Perception
- McGurk Effect – Vision plays a huge role in how we perceive sound,
Aphasia
- Inability to have language
- Wernicke
- Broca – speech production problems, stilted speech, simple sentence but have good comprehension and speech perception
- Wernicke – nonsensical sentences, clear words but without meaning, little comprehension and understanding of speech perception
Babies and Music
- Processing music is innate and babies who are very young can differenitate preffered tunes
- Babies prefer their mother’s singing to other female voices
- consistently prefer consonant to dissonant melodies from as young as 4 months