Lecture 4: Gap Dynamics Flashcards
In the 1930’s Aubreville proposed ___
‘gap mosaic’ model in which the forest cycles through 3 main stages
3 main stages of ‘gap mosaic’ model
1) gap creation: immediately following a tree fall or gap opening
2) Building or regeneration phase: which is characterised by competition for light & other resources leading to regeneration & growth
3) Mature phase: where the high emergent forest canopy is re-establsihed
Gaps are a ___ part of forests dynamics
natural
Forests are a mosaic of __ &__ at different stages of ____
gaps & trees and different stages of regeneration
how are gaps created?
- rain forest trees will eventually die and fall, they may be uprooted or snap
- trees may be blown down
- struck by lightning& fall
how does one tree falling cause so much destruction:
- when one tree falls it often brings many more as crowns are meshed together by lianas
- Large branches continually break off from trees and fall
- Gaps vary in size
Damage to vegetation below varies with
gap size, composition of surrounding vegetation, type of forest
gap fraction is defined as
the fraction of sky visible through the canopy
Gap characteristics of different rain forest types ___
VARY
- Gap size & Gap fraction
gaps create ___ microclimates affecting _______
diverse microclimates affecting light, moisture, temperature & wind conditions
__ is the most limiting factor in the understorey of rain forests
Light
full sunlight is approximately ___ micromoles/m^2/s on a sunny day
2000 micromoles/m^2/s
At la Selva, Costa Rica gaps of approximately 300m^2 experience ___% of full sunlight compared to the understory which received ___ full sunlight
gaps = 8.6-24% understory = 0.4-2.4%
Quality of ___ also changes with an increase in Photosynthetically Active Radiation (PAR) (400-700nm)
LIGHT
Gaps receive differing levels of irradiance from…
full sun in the centre to shade at the edge
Vapour Pressure Deficit, or VPD, is the
difference (deficit) between the amount of moisture in the air and how much moisture the air can hold when it is saturated
once a gap opens competition for ___ & ____ is intense &____ take places
once a gap opens competition for light and nutrients is intense and rapid growth and reproduction take place
there are different stages of succession within gaps:
-rapid colonisation by shrubs, vines, lianas & seedlings of PIONEER TREE species
-rapid growth of short lived light-daminding species which form canopy over 10-30years
-Slower growing shade tolerant (CLIMAX TRESS) species grow in biomass and species richness below the pioneer canopy & become taller. This phase transitions into the next phase when
-shorter lived species die & the more shade tolerant climax trees emerge & re-establish the tall canopy (can take 75-100 years )
HOWEVER LOT OF VARIATION IN THESE PATTERNS
classification for pioneer and climax species:
- broad categorisation based on plant characteristics (functional traits) that influence growth & survival strategies
- pioneer-climax classification represent extremes on a continuum
pioneer species typically produce:
lots of small seeds
seeds of pioneer species are dispersed __
widely –> wind / animals