Plant Bio Exam 1 Flashcards
Herbaceous tissue of the primary plant body develops from the what?
apical meristem
Vascular tissues occurs in what in the stem?
distinct bundles
In woody species, what are produced in the stem & root from activity of other meristems?
secondary tissues
produces wood containing secondary xylem
vascular cambium
produces bark containing secondary phloem and cork
cork cambium
Vascular cambium and cork cambium constitute the plants what?
secondary body
Woody plants are a combination of what?
primary and secondary tissue
A herbaceous plant’s conducting capacity is set after what?
a portion of stem or root is mature
Woody plants become wider every year by what?
accumulation of wood and bark, giving them a greater conducting capacity
Disadvantages to secondary growth (being woody)
- greater need for defenses, both structural and chemical to survive for a long time
- must use energy & nutrient resources for winterizing their bodies in temperate climates
- expensive metabolically to construct wood and bark
- Woody plants may not reproduce until they are several years old due to the energy spent on other activities
All woody trees and shrubs (including gymnosperms) descended from one group of what?
ancestral woody plants that arose about 370 million years ago
- therefore ‘Wood’ is an ancient trait and has evolved infrequently
Most evidence indicates that the first flowering plants were what?
woody
What is the derived condition of the herbaceous condition?
loss of wood
True secondary growth (wood) occurs in what?
- many eudicots
- most basal angiosperms
- all gymnosperms
- but never in ferns or monocots
What initially evolved from a woody ancestor but later lost the ability to produce woody tissues?
monocots
the meristem that produces the wood of the secondary plant body
Vascular cambium
-It originates as a layer of cell that lies between the xylem
and phloem of a vascular bundle in a plant stem
- In herbaceous plants these cells do not divide (usually)
- But in woody plants this region becomes meristematic
Vascular cambium
occur between xylem and phloem within a vascular bundle
Fascicular cells
occur between vascular bundles
Interfascicular cells
To form a complete vascular cambium that completely encircles the plant stem, what two region of cells must become meristematic?
1) Fascicular cells
2) Interfascicular cells
Once developed, the vascular cambium will consist of what?
a single cell layer encircling the stem that is capable of continued division
What two types of cells does the vascular cambium contain?
1) Fusiform initials
2) Ray initials
long and narrow vascular cambium cells
fusiform initials
short, cuboid vascular cambium cells
Ray initials
- long, tapered cells
- divide to produce (to the interior) the elongate cells of xylem (wood): tracheids, vessel elements, fibers, parenchyma
- also divide to produce (to the exterior) elongate cells of phloem: sieve cells, sieve tube members, companion cells, fibers, parenchyma
Fusiform initials
Fusiform initials of the vascular cambium divide longitudinally with what to produce two elongate cells?
periclinal wall (parallel to the meristem)
- one cell remains a fusiform initial (of the meristem)
- the other differentiates into either a cell of secondary xylem or secondary phloem
What (produced to the interior) increases greatly in diameter in growth, pushing the vascular cambia cells outward?
Secondary xylem cells
- produces stress on the cambial cells that can’t keep up
Vascular cambium cells must occasionally divide longitudinally by what (perpendicular to the cambium’s surface)?
anticlinal walls
- allows the cambium to add cells and increase in diameter and keep up with the increasing girth of the woody layer underneath
- short and cube-shaped
- divide to form xylem or phloem parenchyma that functions in storage or as albuminous cells (in gymnosperms)
Ray initials
Within the vascular cambium, Fusiform initials may occur (depending on the tree species):
- in regular horizontal rows (storied cambium)
- irregularly, w/o any horizontal pattern (nonstoried cambium)
Within the vascular cambium, Ray initials are grouped together in what?
short vertical rows
- one cell wide (uniseriate)
- two cells wide (biseriate)
- many cells wide (multiseriate)
Vascular cambium never has what?
large regions of just fusiform initials or just ray initials; it is always mixed
The overall ratio of fusiform initials to ray initials for a species is what?
relatively constant
- under precise genetic/developmental control
Types of wood cells:
- secondary xylem (wood)
- an axial (vertical) system
- a radial (horizontal) system
contains all of the cell types that occur in primary xylem
Secondary xylem (wood)
derived from fusiform initials
Axial (vertical) systems
develops from the ray initials
radial (horizontal) system
The axial system contains what?
- Tracheary elements
- fibers
- parenchyma
tracheids and vessels that carry out vertical conduction of water through the wood
Tracheary elements
provide strength
fibers
serves as a temporary reservoir of water
parenchyma
Hardwoods vs softwoods
- Hardwoods contain large amounts of fibers
- Softwoods contain few or no fibers
Most gymnosperms (softwoods) contain only what?
tracheids in their axial systems
- fibers and parenchyma cells are sparse or absent
In woody angiosperms, the radial system (‘rays’) contains only what?
parenchyma
- arranged in uniserate, biseriate, or multiseriate arrays-
For woody plants in temperate regions: Where do growth rings occur?
in wood due to the differential growth of early (spring) wood versus late (summer) wood
what has high proportions of wide vessels or tracheids?
spring wood
what has fewer vessels or narrower, thick-walled tracheids?
summer wood
What makes up 1 year’s growth, or annual (growth) ring?
Early wood and late wood
wood with vessels found mostly in early wood
ring porous
Ring porous species include:
- oaks
- hickories
- ashes
wood with vessels found throughout
diffuse porous
diffuse porous species include
- red maple
- black gum
- center of a log
- darker, drier, andmore fragrant
heartwood
-outer log
- light colored
Sapwood
- denser
- contains less water
- more aromatic
- better acoustic properties
Heartwood
Heartwood forms when:
- tracheary elements of older portions of wood no longer function in water transport
- tree seals off this old vascular tissue in the wood to avoid fungal hyphae or bacteria from invading and causing rot
What forms inside the old tracheary elements?
tylosis
intruding plug from an adjacent (living) parenchyma cell
tylosis
What produces compounds that inhibit growth of bacteria and fungi?
Xylem parenchyma cells (XP)
- makes cells dark and aromatic
Eventually, the tracheary elements are completely plugged up and filled with what?
defense compounds
When the parenchyma cells die (in heartwood), they leave behind what?
dark, highly decay-resistant cells
What contributes to heartwood?
parenchyma and tracheary elements
In laterally spreading branches, what is necessary to provide support? What is the resulting wood called?
- differential wood strength
- reaction wood
In angiosperms, additional growth occurs mostly on which side of a branch?
upper side; tension wood (pulling up)
- enriched with gelatinous fibers enriched with cellulose
- grows on upper side of a branch
- thicker annual rings on top of branch provide additional strength
- thinner annual rings occur on bottom of branch
Tension wood