Mollusca Flashcards
phylum mollusca
second most species-rich phylum.
-mostly marine, some freshwater & terrestrial.
9 common to most mollusca
- small eucoelom + large haemocoelic circulatory system
- chambered heart
- dorsal body wall with thick epidermal layer = mantle
- glands in mantles create calcareous shell
- ventral body wall muscles develop as muscular foot for crawling, digging
- radula for feeding - toothed, rasping
- complex gut
- gas-exchange systems - ctenidia.
- excretory system includes metanephridia
two types of fluid-filled cavities within the mesoderm
small eucoelom - arises via schizocoely.
hemocoel extensive - blood pumped through short vessels that open into spongy tissues. “open” circulatory system & hydraulic skeleton
2 main clades of extant mollusca
aculifera
conchifera
aculifera - 3 classes within
- small calcareous sclerites into mantle.
polyplacophora
caudofoveata
solenogastres
conchifera - 5 classes within
- most of body covered in one or two large shells
monoplacophora gastropoda bivalvia cephalopoda scaphopoda
mollusc: shells - what order doesnt have shell?
what are three layers?
aplacophoran doesnt have shell. (others may have lost independently)
- periostracum (tough protein) - prismatic (chalky calcareous layer - thickest)
- nacreous layer (shiny, smooth inner layer)
how do pearls form?
pearls form when irritant gets into mantle of a valve, secretes nacre around it to wall it off
mollusc: ctenidia - structures
multi-leaved gills in mantle cavity.
long flattened central axis bearing two haemolymph vessels & ciliated lamellae
mollusc: ctenidia - principle function
gas-exchange: deoxygenated flows thru efferent vessel, rough lamellae where they pcik up O2, back to body cavity via afferent vessel.
– also used for collecting food particles
mollusc :sensory organs (4)
eyes: evolved independently in different lineages; absent, simple, image-forming
statocyst: in conchifera
- tentacles: slim extensions used for touch + chemoreception
- ospharidium: path of chemosensory tissue, near gills
mollusc: reproduction
highly variable.
- mostly separate sexes, sometimes hermaphroditism.
- dertilization mostly external except cephalopoda, gastropoda.
- development indirect with trocophore (sometimes veliger) larva
aculifera
- clade aplacophora = 2 classes
- 3 classes in aculifera
caudofoveata (aplac)
solenogastres (aplac)
lat, polyplacophora = aplacophora
featuers of aplacophora
completely lack shell plates. pointy calcareous spicules in their mantles. small, marine, benthic, burrowers, lack eyes & metanephridia
class caudofoveata - clade?
aplacophora
- no foot.
- do have radula. one pair of ctenidia. no visceral hump. separate sexes
class solenogastres - clade?
no ctenidia, some species lack radula.
- small foot that can be retracted. sequentially hermaphroditic. trocophore
class polyplacophora - clade? common name? defining feature; other features.
clade: aculifera
common name: chitons
defining feature: 7-8 flat dorsal shell plates.
8 pedal retractors, many ctenidia. poor head, lack eye. powerful foot.
repro in polyplacophora
sexes separate, fertilization external.
egg shed in gelatinous string.
indirect development with trocophore larva
conchifera - 4 shared features
shell univalve
shell of basically three layers
statocysts
viscera concentrated dorsally.
class monoplacophora - clade?
- known from?
- mollusc features?
clade conchifera. known from fossils from cambrian. lack eyes, have foot , radula, 3-7 metanephridia ,3-6 pairs of ctenidia