Lecture 4: Origin of Prokaryotes and Eukaryotes Flashcards
Rocks of suitable age contain three types of fossil evidence for prokaryote life:
1) Fossil Stromatolites
2) Fossil microorganisms
- silicified in cherts
- resistant cell walls called acritarchs preserved in siltstones
3) Carbonaceous matter that can be identified chemically as the product of ancient life
Fossil stromatolites:
cyanobacteria on surface & anaerobic bacteria in centre
1) sediment settles
2) live substances rise to surface
3) etc keeps going
cherts
silica grown as nodules
where are siltstones positioned?
bottom of ocean
Prokaryotes dominated the planet for
some 2 billion years
Prokaryotes remain..
relatively unchanged to the present day
by 3,500 Ma we have…
advanced prokaryote life in the oceans. Consisted on complex and diverse communities of stromatolite forming communities on the floors of shallow seas & free living prokaryotes floating around in the oceans.
Both communities include cyanobacteria that produce oxygen as a biproduct of their metabolism.
The oxygen as a biproduct from the cyanobacteria means the-
environment becomes aerobic
Evidence for the earth becoming aerobic:
- banded iron formations
- pyritic conglomerates
- red beds
BIFs
Banded Iron Formations
Early Oxygen Sinks:
1) Volcanic gases that readily combine with O2 (N–NO & NO2; H–H2O; CH4—CO2; H2S–SO4)
2) Dissolved iron that scavenged O2 to form BIFs
3) Microorganisms carrying out aerobic respiration (O2 + organic matter = H2O + CO2 + cellular energy). The earliest organisms were no doubt facultative, respiring aerobically when O2 was available, but switching back to anaerobic fermentation when it was in short supply
2 consideration for considering the origin of eukaryotes
- when were conditions suitable for the origin of eukaryotes
- fossil evidence for eukaryotes
by 2 billion years ago some acritarchs are large enough to,,
suggest that they were Eukaryotes
2 theories of how eukaryotic cell originated:
1) Symbiosis
2) Elaboration of cell membrane
Symbiosis steps:
1) Thermoplasma-like Archaebacterium (nucleosome-like containing chromosome) is engulfed by photo-eukaryote cell = Eukaryotic cell