Lecture 4 Flashcards
Macroevolution
-evolution above the species level: includes big patterns of evolution such as those seen in the fossil record
Microevolution
-evolution within species and populations. Darwinian Natural Selection allows the study of microevolutionary mechanisms that underlie macroevolution
Natural Selection
-process by which the variants of organisms in a population that are best adapted to the environment increase in frequency relative to less adapted variants over a number of generations
Acceptance of natural selection
- not liked in 19th century
- mendelian gentics put down Lamarckian evolution and was consistent with selection
- Early 20th century biologists thought mutation was more important than selection
- 1930s-1940s population genetics shows role of selection, the “Modern Synthesis”
- Modern experiments directly show selection in action in living organisms
Darwin’s 4 postulates
- basis of Natural selectin
- Individuals in a species are variable
- some variation is heritable
- more offspring are produced in each generation than can survive
- survival and reproduction are not random with respect to variation among individuals
Ugly facts about nature
- Darwin recognizes that every environment is full of competition, predation, and death
- potential for selection is very strong
Selective pressures
- birth—–>reproduction
- birth defects, disease, accident, parasites, predation, starvation, competition for mates, environmental extremes
Selection
- normalizing: keeps population at present mean
- directional: change in favored state of population
- sorts variation and does not respond to environment: non random
- acts only on existing traits and depends on what genetic variability is available
- can’t achieve perfection in design
Selection requires
- reproduction
- heredity
- variation in characteristics among individuals
- variation in fitness
fitness
-probability that an individual will survive to reproduce
origin of variation
-not understood at all by Darwin and was a weakness of selection theory until genetics re-discovered
Mendel
- provided basic rules of heredity
- genes as particulate heritable units
Timing of evolution
-evolutionary events may cover millions of years but the action must take place in each generation of an evolving lineage
John Endler
- guppies
- sexual selection: males have bright blue spots
- predation: fish eat brightest guppies first so select for males with small spots
Origin of Variation
- to be heritable, variation has to result from a genetic change in DNA-a mutation
- mutations occur without regard for the external environment or needs of the organism
- mutations have to change the phenotype to be subject to selection
- there are silent mutations aplenty. they accumulate in the genome=non-Darwinian evolution