Lecture 11 - The Other Senses Flashcards
Audition stimulus
- sound waves
- pitch
- loudness
- timbre
Sound waves
vibrations of an object cause air particles to vibrate (oscillations of compressed and rarefied air). (like ripples in water)
Pitch
frequency of vibration (Hz).
Loudness
intensity (more vigorous vibrations = more intense stimuli).
Timbre
complexity of sound (mixture of vibrations).
- could tell even though they are same notes they’re coming from 2 diff instruments
More waves per sec, the ____ the pitch
HIGHER
Anatomy of the ear
- sound funneled through ear canal to ear drum (tympanic membrane).
- middle ear – bones set to vibrate by ear drum.
- malleus (hammer), incus (anvil), stapes (stirrup). - connect to cochlea
- cochlea – energy transmission from middle ear to inner ear (from air to liquid).
sound waves –> mech energy –> energy in inner ear filled with liquid (physical waves of liq) going to cochlea
Auditory pathway dets
- cochlear nerve (branch of vestibulo-cochlear).
- 95% synapse with inner hair cells.
- outer hair cells are EFFECTOR cells that alter effects of vibrations on inner cells.
(depending on which hair cells in cochlea get stimuluated, we perceive diff sounds)
Auditory cortex – primary pathway
Auditory Nerve
- -> Cochlear Nuclei
- -> Superior Olivary Nucleus
- -> Auditory Cortex
- -> Medial Geniculate Nucleus
- -> Inferior Colliculus
- CONTRAlateral and IPSIlateral projections – information from the cochlear goes to both hemispheres.
- TONOTOPIC organization. - depending on vibration freq, diff parts of basilar mem. will be stimulated & vibrate – hair cells will in turn be excited & send signals to brain
Perception of pitch
PLACE CODING – DIFFERENT PARTS of basilar membrane FLEX in respond to DIFFERENT FREQUENCIES – point of maximum vibration is tightly coupled to frequency.
Perception of loudness
loudness transmitted to CNS by rate of firing (louder sounds – higher firing rate).
loudness of low frequencies signaled by number of neurons firing (because rate changes pitch in these neurons).
Perception of timbre
same note on different instruments – timbre gives the different qualities.
- get richness of music b/c of timbre
Perception of spatial location
PHASE DIFFERENCE – the difference in arrival times of sound waves at each of the eardrums. Left/right judgments are more accurate (b/c we use a race to the sup. olivar nucleus 1st) than front/back (hits at same time).
LOCATION BY INTENSITY – intensities are absorbed by the head creating a “sonic shadow”.
timing & intensity help localize sound
Association cortex and pattern recognition
What vs Where. Brain activity in response to judgments of category (red) and location (blue) of sounds.
IFG = inferior frontal gyrus, IPL = inferior parietal lobule, MFG = middle frontal gyrus, SFG = superior frontal gyrus, SPL = superior parietal lobule.
Vestibular system
semicircular canals – angular acceleration (rotation of the head). (when you move, they move - can tell direction of which way you turned your head with your eyes close)
vestibular sacs – responds to gravity.
Vestibular functions include
balance
head position
eye / head compensation (vestibulo ocular reflex)
Vestibular system
Low frequency stimulation of the vestibular sacs can cause…
NAUSEA
stimulation of canals can cause dizziness.
Vestibular system
nystagmus
– rhythmic left/right eye movements.
caloric stimulation (cold water in the ear) causes nystagmus (twitching)
Vestibular anatomy
canals approximate head planes (sagittal, transverse and horizontal).
most axons synapse on vestibular nuclei in medulla.
some direct synapses with cerebellum.
cortical connections – dizziness.
lower brain stem connections – nausea & vomiting.
Over-representation in the Motor and Somatosensory strips
Regions requiring more fine motor control (or with greater density of tactile receptors) will have larger representations in the cortex
Cutaneous senses
touch (skin receptors).
Kinesthesia
body position and movements (joints, muscles, tendons).
Organic senses
receptors in and around internal organs (butterflies in the stomach).
The Stimuli
Touch
pressure – deformation of the skin (pressing down on arm)
vibration – surface texture (touching diff senses like sand paper)
pain – tissue damage
The Stimuli
Kinesthesia
stretch receptors in muscles (info re: muscle length)
stretch receptors in tendons (info re: force)
joint receptors provide info re magnitude and direction
Skin – a vital organ!
protection (wound is opening for pathogen)
thermoregulation (sweat or restriction of circulation)
Pressing on skin =
indentation (Ruffini)
Rubbing fingers on a surface & feeling vibrations =
vibration (Pacinian)
Glabrous skin
glabrous – hairless, more complex mixture of free nerve endings and axons
- palms & bottom of feet
- good at picking up MINUTE DETS b/c of all the nerve endings they have small receptive fields
2-point test
- prongs vary on test
- you can go v. close together on fingertips for ex & still determine that is 2 separate pricks
Pacinian corpuscle
sensitive to vibration.