leaves Flashcards
three ecological types of plants
mesophytes
xerophytes
hydrophytes
mesophytes
require moderate amounts of water
xerophytes
adapted to low levels of water
hydrophytes
adapted to high levels of water
eudicots leaves in sun vs shade
- in sun leaves have denser palisade
-shade have thinner less dense palisade
ex of phenotypic plasticity
major veins
- may have a bundle sheath of parenchyma (largest do not)
- these veins have less exit/entry from the mesophyll
- serve more as a transport function in/out of leaf
- sometimes may undergo limited amount of secondary growth
- also may have bundle extensions of collenchyma for support
- some species move water to the epidermis by bundle extensions
minor veins
- often lack bundle extensions
- always have a bundle sheath with few chloroplasts
- little space between bundle sheath cells
- helps control exit/entry to veins
- large degree of control because of large number of minor veins
netted (reticulate) venation
in a typical eudicot leaf
-mesophyll divided into subregions called areoles
a paradermal secttion
- dense packing of the palisade mesophyll
- vein network
- more air spaces in spongy mesophyll
- stomatal density of adaxial and abaxial differs
vascular tissue
- bundle sheath is part of vascular tissue
- xylem and phloem similar in c3 and c4 grass leaves
- larger veins resemble stem bundles - have vessel elements, tracheids, parenchyma and a protoxylem lacuna
- minor veins generally have few tracheids, and no vessels
- sieve tube members and companion cells phloem
evergreen angiosperm leaves
- some woody eudicots retain leaves year round
- called broad-leaved evergreens
- grow in low nutrient status habitats
- slow growers and stress tolerators
- plants conserve organs as some nutrients not salvageable but must be frost hardy and xerophytic (cold causes xerophytic things )
- tolerant of ice crystal formation within tissues, not cells (a tolerance of all temperate woody species)
- particularly found in ericaceae
- ericoid describes general leaf morphology
- example of convergent evolution
different leaf habitat
stays on plant of the winter
- leaf said to be wintergreen = not true evergreen
- turns red/purple in late fall
- leaves senesce in spring
leaf abscission
- controlled shedding of leaves
- important in woody species ie. perennials
- occurs yearly in deciduous species
- or at longer intervals for evergreens
- regulated by hormones
- leaves drop in response to environmental cues
steps of leaf abscission
- environmental signal
- retrieval
- abscission layer formation
- protective layer formation
- vascular disconnection
environmental signal
Photoperiod more reliable
Temperature can also influence
Warm temperatures in fall can inhibit onset, especially in non-native species
retrieval
Plant recovers as many soluble nutrients as possible eg. Mg+2, sugars, amino acids
Stores in parenchyma in stem
Some minerals not mobile
ie. not retrievable eg. Ca+2
abscission layer formation
New thin-walled parenchyma cells form at base of petiole
From dedifferentiating parenchyma
Cell walls of pre-existing cells also begin breaking down here
Forms an even break line
protective layer formation
Another layer forms to the inside of the abscission layer
Is suberized to prevent or minimize water loss and pathogen entry
Creates a leaf scar on the stem
vascular disconnection
Vessels in midrib of petiole still open Leaf may hang on for a time Nearby xylem parenchyma produce outgrowths that enter pits of vessel Is to seal off vessels Outgrowths called tyloses Can see bundle scars within leaf scars
fall colors in deciduous leaves
A shorter photoperiod and cooler temperatures initiate leaf fall
Chlorophyll broken down by light all summer, but is not replenished in fall
Fat soluble carotenoids more resistant
In some species, water soluble anthocyanins are produced de novo in vacuole
Other plants always contain anthocyanins
Anthocyanins are increased with cold nights and sunny days
morphology
- major ps organ
- important for other metabolic functions like synthesis of amino acids and secondary metabolites
- most variable organ in form and anatomy
- different environments have differing selection pressures
blade/lamina
-flattened portion