Learning and Ethology Flashcards
Shaping, also known as Differential Reinforcement
Operant conditioning phenomenon in which you reinforce only the behavior (response) you wish to increase and extinguish all other behaviors (responses). For example, suppose you want to train your dog to fetch your slippers. So you reinforce successively closer approximations to the desired behavior. You might begin by reinforcing your dog every time she looks at your slippers. Once she’s doing that consistently, you stop reinforcing her for just looking at your slippers (i.e. you extinguish that behavior) and reinforce her only when she walks toward your slippers. And so on…
The 5 consequences of operant conditioning,
as specified by B. F. Skinner
- Positive reinforcement: occurs when a behavior (response) is rewarding, or the behavior is followed by another stimulus that is rewarding, thus increasing the frequency of the behavior.
- Negative reinforcement: subtype escape occurs when a behavior (response) is followed by the removal of an aversive stimulus; subtype avoidance occurs when an organism gets a warning that an aversive stimulus will soon occur, and the appropriate response completely avoids the aversive stimulus.
- Positive punishment, or punishment by contingent stimulation: occurs when a behavior (response) is followed by an aversive stimulus.
- Negative punishment, or punishment by contingent withdrawal: occurs when a behavior (response) is followed by the removal of a rewarding stimulus.
- Extinction: occurs when a behavior (response) that had previously been reinforced is no longer reinforced; the behavior eventually becomes extinguished.
Acquisition
“Acquisition” is a term used in both classical and operant conditioning. In classical conditioning, it refers to the period during which an organism is learning to associate the CS with the UCS.
Ivan Pavlov
Usually credited for the basic principles of classical conditioning. He noticed that through experience, stimuli that previously had no relation to a specific reflex could come to trigger that reflex. A reflex is simply an unlearned response that is elicited by a specific stimulus. The idea of classical conditioning is to cause a neutral stimulus (for example, the ringing of a bell) to become paired, or associated, with a stimulus that elicits a reflex (e.g., food powder placed into dog’s mouths). For classical conditioning to be successful, the neutral stimulus, or conditioned stimulus, must be presented before the unconditioned stimulus (the food powder). Presenting the CS before the UCS is known as “forward conditioning.”
Karl von Frisch
(1886 - 1982) Austrian ethologist who received the Nobel Prize in Physiology or Medicine in 1973, along w/ Nikolaas “Niko” Tinbergen and Konrad Lorenz, for their discoveries concerning organization and elicitation of individual and social behavior patterns in animals. Much of his personal work centered on investigations of the sensory perceptions of the honey bee. Von Frisch and others found that honey bees are able to communicate the direction and the distance of a food source to their fellow hive members by means of special movement patterns, often called “dances.”
Ethology, past and present
As opposed to early ethologists, modern ethologists tend to deemphasize the instinctual bases of behavior and focus more on the question of why the animal behaves as it does and not in some other manner. To answer this question, ethologists attempt to discover the evolutionary significance of behavior.
Darwin’s Theory of Natural Selection,
in 3 steps
- There are genetic differences between members of a species.
- If a specific genetic variation increases the chances of reproduction, it will tend to get passed down to the next generation. If a specific genetic variation decreases the chances of reproduction, it will tend not to get passed down to the next generation.
- Over time, more and more members of the species will tend to have the genetic variations that increase their chances of reproduction and less and less of the species will tend to have the genetic variations that decrease their chance of reproduction.
Reproductive Fitness
(also called Individual Fitness or Personal Fitness)
vs.
Inclusive Fitness
Reproductive fitness means the number of offspring that live to be old enough to reproduce. If the currency of natural selection is reproductive fitness, then altruism (in this context, behavior that increases the reproductive fitness of other members of one’s species while decreasing one’s own) is difficult to understand. By contrast, the theory of kin selection suggests that animals act to increase their inclusive fitness, rather than their reproductive fitness. Inclusive fitness takes into account not only the number of offspring who survive to reproductive age, but also the number of relatives who survive to reproductive age. If the currency of natural selection is inclusive fitness, then altruism makes sense.
Sociobiology
A field of biology that aims to examine and explain social behavior in terms of evolution. Sociobiology draws from the disciplines of psychology, ethology, anthropology, evolution, zoology, archaeology, and population genetics. The scientist most associated w/ sociobiology is E. O. Wilson. Wilson is adamant in his belief that behavior is due to a complex and dynamic interplay between genetics and the environment. Wilson’s biological specialty is myrmecology, or the study of ants.
Fixed-Action Patterns (FAPs)
When you observe the behavior of animals in their natural habitat, you see that certain action patterns are relatively stereotyped and appear to be species-typical. These actions are called fixed-action patterns (FAPs). Because they are considered innate, it is perhaps easy to confuse a FAP w/ a Pavlovian unconditioned response, which is also considered to be innate. The difference is that a FAP tends to be more complex than a Pavlovian unconditioned response. So, for example, an unconditioned response might be something like salivation or an eye-blink whereas a FAP might be something like rolling an egg back to a nest, or a species-typical courtship ritual.
Nikolaas “Niko” Tinbergen
(1907 - 1988) Dutch biologist, ornithologist, and ethologist who shared the 1973 Nobel Prize in Physiology or Medicine with Karl von Frisch and Konrad Lorenz for their discoveries concerning organization and elicitation of individual and social behavior patterns in animals. He is regarded as one of the founders of modern ethology, and was one of those ethologists who introduced experimental methods into the field, enabling the construction of controlled conditions outside the laboratory. Tinbergen also conducted one of the most famous experiments in ethology–on aggression in male sticklebacks (a fish).
Sign Stimuli and Releasers
Sign stimuli are features of a stimulus that are sufficient to bring about a particular FAP. Releasers are sign stimuli that function as signals from one animal to another. However, the two terms are often used interchangeably. In general, a specific FAP will be elicited by only one sign stimulus or releaser.
Supernormal Stimulus
In his experiment on aggression in male sticklebacks, Tinbergen was able to design a model that elicited the aggressive response more reliably than an actual male stickleback. A stimulus that is more effective at triggering the FAP than the actual stimulus found in nature is called a supernormal stimulus.
Garcia Effect
Conditioned taste or smell aversion associated w/ a negative reaction, such as nausea or vomiting. Discovered by John Garcia via a classical conditioning experiment w/ thirsty rats. The experiment was a 2x2 factorial design involving two unconditioned stimuli (a shock or a nauseating drug) and two conditioned stimuli (sweet water or bright-noisy water). Conditioning was successful only in two of the four conditions: when the nauseating drug was paired with the sweet water, and when the shock was paired with the bright-noisy water. These results are explained by a phenomenon called preparedness. Rats seem to have an inborn, biological tendency to associate illness w/ something they ingested and to pair sights and sounds w/ externally induced pain. Humans also are prone to associating illness with what they ingest.
John Garcia
Studied taste-aversion learning and proposed that some species are biologically prepared to learn connections between certain stimuli.
Biological Constraints
To challenge behaviorist theories, researchers point to evidence that different species have different inborn predispositions for how and what they are able to learn. These predispositions are called biological constraints, and they affect both classical and operant conditioning. One exaple is the Garcia effect, which illustrates the principle of preparedness and involves classical conditioning. Another example is instinctual drift, which can interfere with attempts to shape behavior through operant conditioning.
Wolfgang Köhler on Problem Solving
Wolfgang Köhler (who cofounded Gestalt psychology) disagreed w/ Edward Thorndike (who conducted experiments w/ cats in puzzle boxes and formulated the law of effect). Thorndike argued that all problem solving is trial-and-error. Köhler argued that given the opportunity, some animals can learn by insight. Insight is the perception of the inner relationships between factors that are essential to solving a problem. Köhler placed chimps in enclosed play areas and suspended food high above (in their sight, but out of reach). Typically, chimps would begin by trying to get the food directly by reaching for it. When that failed, they would typically stop and survey the situation. After a period of time, and often suddenly, they would try a different approach based on a novel way of using items in the cage.
Conditioned Aversion
Behavior therapy based on classical conditioning. May be used to help, among others, people with alcohol problems, people addicted to smoking, and people with problematic paraphilias. In conditioned aversion, the stimulus that attracts the client becomes paired w/ an aversive unconditioned stimulus associated with punishment. Negative feelings become associated with the undesirable behavior and the client stops being attracted to the behavior. In technical terms, conditioned aversion involves pairing a desired (but problematic) CS with an aversive UCS.
Nikolaas “Niko” Tinbergen’s
famous experiment on aggression
in male sticklebacks
The stickleback is a fish whose males establish territories during the spring breeding season. If a male swims into the territory of another male, he’s likely to be attacked. Tinbergen and his associates found that the red belly of the invading stickleback was the most important element in triggering the aggressive behavior, since even a crude model with the belly painted red was apt to be attacked. Therefore, a red belly is a sign stimulus, and also a releaser, which triggers aggression in male sticklebacks during the spring.
Innate Releasing Mechanism (IRM)
The FAP follows automatically once the organism perceives the sign stimulus. In fact, even if the stimulus is removed in the middle of the behavioral sequence, the animal will continue to perform the actions as if the stimulus was still present. Because of this, ethologists have suggested that there must be some mechanism in the animal’s nervous system that serves to connect the stimulus with the right response. They call this mechanism an innate releasing mechanism (IRM).
Reproductive Isolating Mechanisms
Reproductive isolating mechanisms are behaviors that prevent animals of one species from attempting to mate with animals of a closely related species. They work by providing an animal with a way of identifying others of its own species. An example is the species-specific call given by male black-headed gulls, enabling female black-headed gulls to find them. Isolating mechanisms are found only in locations where closely related species share a common environment.
Observational Learning
Albert Bandura demonstrated by his classic “Bobo doll” experiment that behavior can be learned by observation, or what he termed vicarious reinforcement. Vicarious reinforcement refers to the process of learning behaviors through observation of reward and punishment, rather than through direct experience.
Keller Breland
and
Marian Breland Bailey
Behavioral psychologists who used shaping in an attempt to train a raccoon to pick up coins and deposit them into a piggy bank. They encountered a difficulty known as instinctual drift: the raccoon kept reverting to a species-specific behavior pattern. The raccoon never fully learned the desired procedure. Instinctual drift is a kind of biological constraint.
How the Garcia effect, i.e. taste-aversion learning,
differs from the “standard” model of classical conditioning
Learned taste aversion can occur after only a single trial. One pairing of the CS and UCS is all it takes. This is unlike most classical conditioning where many trials are required for the conditioning to fully develop. What’s more, taste-aversion learning can take place even if the UCS occurs up to 24 hours after the CS. Usually, the optimal time period between the CS and UCS is several seconds after the CS. But typically, if you become ill up to 24 hours after eating a new food, you will probably be averse to eating any more of that food.