Exam 2 Flashcards
Ecological Species Concept
defines a species by its ecological niche; applies to sexual and asexual species and emphasizes the role of disruptive selection
Macroevolution
-broad patterns of evolutionary change above the species level
-cumulative effect of many speciation and extinction events
Speciation
the process by which one species splits into 2 species
What does speciation contribute to biology?
-produced tremendous diversity of life
-helps to explain the unity of life
-forms a conceptual bridge between micro- and macroevolution
Temporal Isolation
species that breed at different times of day, in different seasons, or different years –can’t mix their gametes
Microevolution
changes in allele frequencies in a population over time
Behavioral Isolation
-prezygotic barrier
-courtship rituals and other behaviors unique to a species are effective barriers to mating
Biological Species Concept
-have the potential to interbreed in nature
-produce viable, fertile offspring
-don’t produce viable, fertile offspring with members of other such groups
Reproductive Isolation
-when biological barriers impede members of 2 species from interbreeding and producing viable, fertile offspring
-limits the formation of hybrids
Habitat Isolation
-prezygotic barrier
-two species that occupy different habitats within same area may encounter each other rarely, if at all
What was early life on Earth like?
-inhospitable
-little to no oxygen in atmosphere
-meteorite impacts and volcanic eruptions common
-seas full of mineral salts
Gametic Isolation
-prezygotic barrier
-sperm of one species may not be able to fertilize eggs of another species
Prezygotic Barriers
-block fertilization from occurring by impeding different species from attempting to mate
-preventing the successful completion of mating
-hindering fertilization if mating is successful
Mechanical Isolation
-prezygotic barrier
-mating attempted, but morphological differences prevent its successful completion
Postzygotic Barriers
-prevent hybrid zygotes from developing into viable, fertile adults
Reduced Hybrid Variability
-genes of different parent species may interact in ways that impair the hybrid’s development or survival in its environment
Reduced Hybrid Fertility
Meiosis may fail to produce normal gametes, resulting in sterility if the parent species have chromosomes of different number or structure (i.e. mules)
Hybrid Breakdown
First generation of hybrids are viable and fertile, but offspring in the next generation are feeble or sterile
What are the limitations of the Biological Species Concept?
-cannot be applied to fossils or asexual organisms (including all prokaryotes) because mating can’t be observed
-emphasizes absence of gene flow, but gene flow occurs between many morphologically and ecologically distinct species (i.e. grizzly bears + polar bears = grolar bears)
Morphological Species Concept
-distinguishes a species by its structural features
-applies to sexual and asexual species, doesn’t require info on the extent of gene flow
-Disadvantage: relies on subjective criteria
Allopatric Speciation
-gene flow restricted between populations by geographic isolation
-intrinsic barriers to reproduction due to genetic change driven by processes including divergent selection and genetic drift
-reproductive barriers prevent interbreeding even if contact restored
Sympatric Speciation
-reproductive barrier isolates subset of populations without geographic separation
-result from polyploidy, sexual selection, or natural selection resulting from switch in food or habitat
What are the two types of polyploidy?
Allopolyploids and Autopolyploids
Autopolyploidy
-have more than 2 sets of chromosomes from a single species
-in plants, mitotic errors can result in production of tetraploid cell from diploid cell
-fertile offspring produced through self-fertilization or mating with other tetraploids
-mating between tetraploids and diploids produces triploid offspring with reduced fertility
Allopolyploidy
-have 2 sets of chromosomes from different species
-don’t pair during meiosis, resulting in hybrid sterility (sterile hybrids can reproduce asexually)
-formed if chromosome number doubles in subsequent generations
-successfully interbreed with each other but not with either parent species
-diploid number of new species = sum of diploid number of both parents
Habitat Differentiation
-sympatric speciation can result from exploitation of new habitats or resources
example: apple maggot flies evolved after switching hosts from hawthorn to apple
Hybrid Zones
-a region where members of different species mate and produce hybrid offspring
-some hybrid zones form as narrow bands where habitats of 2 or more closely related species meet
-often occurs as isolated patterns scattered across landscape rather than continuous band
How does environmental change affect hybrid zones?
-changing environmental conditions can cause them to relocate
-can also drive production of new hybrid zones
-alleles can be transferred from one parent species to the other through breeding between parents and hybrids (may help parent species cope with changing environments)
What happens if hybrids don’t become reproductively isolated from their parent species?
Reinforcement, Fusion, or Stability
Reinforcement
-strengthening reproductive barriers
-if hybrids are less fit, strong selection for prezygotic barriers should reduce hybrid population
-should be stronger for sympatric than allopatric populations
Fusion
-weakening reproductive barriers
-there can be substantial gene flow between species if hybrids are as fit as parents
-reproductive barriers can weaken and 2 parent species may fuse into a single species
Stability
-continued formation of hybrid individuals
-extensive gene flow from outside hybrid zone can overwhelm selection for an increase in reproductive isolation inside hybrid zone
Punctuated Equilibria
-periods of apparent stasis punctuated by sudden change found in the fossil record
-other species appear to have changed gradually over time
Average time of speciation
6.5 million years
What do all cells have?
-enzymes (carry out reactions)
-plasma membrane
-genome of DNA
Protocell
-believed to be predecessor of cells
-membranous sac containing interacting organic molecules
What was the dynamic between RNA and DNA originally?
-RNA may have preceded DNA
-RNA was THE genetic material and dominant
-switching from RNA to DNA would’ve made genome more stable
Ribozymes
-RNAs that act like enzymes
-support the idea of RNA being dominant genetic material at first
Cyanobacteria
-evolutionary bacteria; first to release oxygen through photosynthesis
-over time, oxygen released by cyanobacteria changed earth’s atmosphere
What did the addition of oxygen do to Earth?
-created the ozone
-favored aerobic life forms
-beginning of ATP-forming metabolic pathway (key innovation in evolution of eukaryotic cells)
First Eukaryotes
-Protists
-proven by their biomarkers and fossils dating back more than 2 billion years
-their diversification led to plants, fungi, and animals
Eukaryotes
-defined by having membrane-enclosed organelles
-some probably evolved from infoldings of plasma membrane in prokaryotic ancestors
-mitochondria and chloroplasts resemble bacteria; maybe evolved through endosymbiosis
Endosymbiosis
-an evolutionary process where one cell enters and survives in another cell
-over generations, host and guest cells begin to depend on each other for essential metabolic processes
-proof: modern protists who have bacterial symbionts inside them
Bacteria
-small cells with DNA and ribosomes
-no nucleus or typical eukaryotic organelles
-most abundant and metabolically diverse organisms
-autotrophs and heterotrophs
-most are unicellular, though some species form colonies
Autotroph
-bacteria that self-feed