2. Hearing Flashcards
Nature of sound:
sound caused by…
That are characterised by…
simplest sound wave is…
air pressure waves
amplitude (dB) - loudness
Frequency (Hz) - pitch
Phase - position within a cycle
a pure tone - sine wave
Nature of sound:
sound caused by…
That are characterised by…
simplest sound wave is…
air pressure waves
amplitude (dB) - loudness
Frequency (Hz) - pitch
Phase - position within a cycle
a pure tone - sine wave
human hearing range is …
20-20,000 Hx
Typical vocal range…
80-1100Hx
Nature of sound:
complec sounds can be built up from…
a series of sine waves with varying amplitudes, frequency and phase
can be decomposed into components by Fourier analysis
Nature of sound:
what is the fundamental?
What are complex sounds made up of?
fundamental = lowest frequency component of a complex sound
many complex sounds are made up of harmonics, - integer multiples of the fundamental
Basic physiology of the ear
components of the OUTER EAR
Pinna
- increases the sound amplitude
- Helps determine the direction from which a sound is coming form
External auditory canal
- provides protection
- increases the sound amplitude
Eardrum (tympanic membrane)
- vibrates in response to sound waves
Moves bones in the middle ear
Basic physiology of the ear
Components of the MIDDLE EAR
ossicles
- Malleus
- Incus
- Stapes
smallest bones of human body
ossicles transmit the vibrations of the eardrum in to the cochlea through lever actions
They also provide protection against high amplitude sounds
- muscles attached to the ossicles restrict the bones’ movements
Basic physiology of the ear
components of the INNER EAR
semicircular canals
- important for vestibular sense (sense of orientation)
Cochlea
- contains auditory sensory receptors
Basic physiology of the ear
The cochlea
- oval window
The cochlear is filled with a watery liquid that moves in response to the vibrations coming from the middle ear
oval window
- membrane covering opening in the cochlea
- the stapes is attached directly to oval window (where the vibrations get into the cochlea
- much smaller than eardrum - size difference further helps amplify sounds waves
Basic physiology of the ear
The cochlea canals
three canals of the cochlear
- vestibular canal
- tympanic canal
- cochlear duct
separated by
- Reissner’s membrane
- basilar membrane (on which hair/auditory receptor cells located in cochlear duct)
Basic physiology of the ear
The cochlea canal and membrane orientation/order
vestibular canal > Reissner's membrane >> cochlear duct >>> Basilar membrane >>>>Tympanic membrane
Basic physiology of the ear
vibrations to neural signals. HOW?
the three membranes vibrate in response to vibrations of the oval window
Wen the basilar membrane vibrates, hair cells are also set in motion
converts the vibrations in to neural signals
the auditory cortex
What auditory tasks can be performed without the auditory cortex present?
- onset of sound
- changes in sound intensity
- Changes in sound frequency
Typical vocal range…
80-1100Hx
Nature of sound:
complec sounds can be built up from…
a series of sine waves with varying amplitudes, frequency and phase
can be decomposed into components by Fourier analysis
Nature of sound:
what is the fundamental?
What are complex sounds made up of?
fundamental = lowest frequency component of a complex sound
many complex sounds are made up of harmonics, - integer multiples of the fundamental
Basic physiology of the ear
components of the OUTER EAR
Pinna
- increases the sound amplitude
- Helps determine the direction from which a sound is coming form
External auditory canal
- provides protection
- increases the sound amplitude
Eardrum (tympanic membrane)
- vibrates in response to sound waves
Moves bones in the middle ear
Basic physiology of the ear
Components of the MIDDLE EAR
ossicles
- Malleus
- Incus
- Stapes
smallest bones of human body
ossicles transmit the vibrations of the eardrum in to the cochlea through lever actions
They also provide protection against high amplitude sounds
- muscles attached to the ossicles restrict the bones’ movements
Basic physiology of the ear
components of the INNER EAR
semicircular canals
- important for vestibular sense (sense of orientation)
Cochlea
- contains auditory sensory receptors
Basic physiology of the ear
The cochlea
- oval window
The cochlear is filled with a watery liquid that moves in response to the vibrations coming from the middle ear
oval window
- membrane covering opening in the cochlea
- the stapes is attached directly to oval window (where the vibrations get into the cochlea
- much smaller than eardrum - size difference further helps amplify sounds waves
Basic physiology of the ear
The cochlea canals
three canals of the cochlear
- vestibular canal
- tympanic canal
- cochlear duct
separated by
- Reissner’s membrane
- basilar membrane (on which hair/auditory receptor cells located in cochlear duct)
Basic physiology of the ear
The cochlea canal and membrane orientation/order
vestibular canal > Reissner's membrane >> cochlear duct >>> Basilar membrane >>>>Tympanic membrane
Basic physiology of the ear
vibrations to neural signals. HOW?
the three membranes vibrate in response to vibrations of the oval window
Wen the basilar membrane vibrates, hair cells are also set in motion
converts the vibrations in to neural signals
Central auditory pathways
nerve fibres from each cochlear synapse in a number of sites on the way to the primary cortex
the pathway is…
The cochlear > The cochlear nucleus >> The superior olivary nucleus >>> The inferior colliculus >>>> The medial geniculate nucleus
NOTE: signal at cochlear nucleus splits and goes to each of the superior olivary nuclei
> beyond this point, input from both ears is present in both hemispheres
auditory space perception
basilar membrane does not directly indicate sound locations so
How is auditory space percived?
it is a biaural process
- interaural time different
»onset difference
» phase difference - interaural intensity difference
the auditory cortex
comparing auditory jask abilities with and without the cortex,
What can be concluded
the cortex deals with more complex auditory tasks while the lower structures deal with simple aspects of sound
speech perception requires structures beyond the primary auditory cortex
theories of encoding
how do auditory system isolate and identify the frequencies of sounds.
HOW?
basilar membrane is 30mm long with varying stiffness and widths along its length.
Traveling waves move along the basilar membrane and peak at different points depending on the frequency of the sound
Thus the location of the peak identifies the frequency of the sound
stimulating auditory nerves at different cochlear locations leads to perception of sound in different pitch
theories of encoding
Who suffers from frequency specific hearing loss?
people who have damage to specific parts of the cochlea
theories of encoding
what are tonotopic maps?
maps present in the auditory system
depict how auditory neurons are arranged in an orderly manner
pitch perception
monoaural encoding refers to…
how sound frequency is encoded in each ear
i.e. cochlear nucleus pathway
pitch perception
biaural pitch encoding refers to…
how structures beyond the cochlear nucleus should be contributing to pitch perception
missing fundamental can be perceived when harmonics are presented in one ear
and can also be receive when presented in different jars
Loudness perception
what are the basic mechanisms of loudness perception
overall firing rate
range of firing
Loudness perception
what factors affect loudness perception
sound duration -
longer=louder
frequency
Loudness perception
how does frequency effect loudness
generally higher frequencies are perceived to be louder
as amplitude increases the effect of frequency becomes smaller
auditory space perception
in auditory space perception, you try to determine a sound’s…
horizontal direction (azimuth)
vertical direction
distance
auditory space perception
describe interaural time diference
unless a sound is directly in front or behind, it reaches two ears at different times (onset difference)
> detected by a simple “delay line” mechanisms in brain
same sound will most likely be in two different phases when it reaches each ear (Phase difference)
> but the phase difference is less useful for localising high-frequency sounds
auditory space perception
describe inter aural intensity diference
the same sound should be a bit more intense at the eat that is closer to the sound source
> the energy of a sound decreases as it travels farther
> the head works as a barrier that reduces the intensity of the sound (sound shadow)
> this effect is more pronounced for high frequency sounds
auditory space perception
How does frequency effect auditory space perception
interaural time difference (phase) is useful for localising low frequency sounds
interaural intensity difference is useful for localising high frequency sounds
auditory space perception
How is head movement useful for auditory localisation
by changing the position of the ears you can experience changes in interaural time/intensity differences