Topic 5 - Fundamentals of Evolution : Drift and Speciation Flashcards
Drift explanation - spandals of San Marco
Architects wanted arches on a rectangular space
These created triangular spaces between the arches and the rectangles
These were evnetally used as sapce for paintings
Did they need space for paintings so the made spandals or did they need arches and spandals were a bi-product?
It was the bi-product!
Same with t rex small arms. Dd evolution make the small arms for a purpose or did it create large legs and a head and so the arms grew smaller to accomodate these and not fall over.
Not all evolution is selection, sometimes it is other things
Genetic Drift
Evolution by chance
Hardy-Weinburg equilibrium
Used to predict what the frequency of genotype will be in the next generation
G is a gene of interest
2 alelles G and g
veryone has 2 alleles
DIfferent combos give different phenotypes
G (p) g(q) female
G(p) GG Gg
g(q) Gg gg
Male
Equation
G (p) 50% g(q) .5 female
G(p) GG Gg .5 g(q) Gg gg .5 Male
p2 + 2pq + q2=1
pxp= .5 x .5 = .25 25% are GG
2(pxq)= 2x (.5 x .5) = .5 50% are Gg
qxq - .5x.5 =.25 25% are qq
Assuming assumptions hold, this is what you would expect in the next generation
Hardy-Weinberg equilibrium assumptions (5 things)
1) No mutation
2) No selection
3) No genetic drift
4) No gene flow (immigration)
5) Random mating (no sexual selection)
ie NO evolutionary change
Why calculate hardy-weinberg equilibrium?
Allows us to study evolution
You can compare the predicted frequency with the actual frequency to determine whether the popultion is evolving
ALso used in medicine to estimate the f of carriers (heterozygotes) for recessive diseases.
EARTHQUAKE!!! (example)
8 individuals, 2 alleles each
f of G = .5
f of g =.5
2xGG
4xGg
2xgg
id 1x Gg and the two gg died, 10 alleles left.
2 GG
3Gg
0gg
Counting them: 10 G and 3 g
f of G = .7
f of g = .3
p2+2pq+q2=1
p2 = .7X.7 = .49 so 50% GG
2pg = 2x(.7x.3)=.42 thus 40% are Gg
q2 + .3x.3 = .0.9 so 10% are gg
This happened by chance. The earthquake did not select them. Tis is the idea behind genetic drift.
Key points for genetic drift
1) effects are random
2) effects are larger in small populations sp they change faster
Two ways genetic drift can result in big changes: 1 - Genetic Bottleneck
An extreme example of genetic drift where an event leads to a big decrease in population size and genetic diversity.
eg a flood where only a few survive, taken at random and y chance (they did not have a survival advantage)
The new population will descend only from this small number of individuals (with their small amount of genetic diversity)
Two ways genetic drift can result in big changes: 1 - Founder effect
A few individuals found a new population somewhere new
eg lizard moves to a new island by chance. All populations are now descended from this founder
There is less genetic diversity and less variation for selection to act on BUT the small population means genetic drift can have a big effect.
Eg lizards move to new island. Random mutation causes bigger claws, descendants climb trees and do well cos its only them up there. Beach lizards random mutation makes them yellow and this protect them from predation
After millions of years, the populations of lizards on the island (descended from the founder that moved there) are totally different from the population on the mainland
A species is
a population whos members can produce offspring that are
(1) viable - survive after birth
(2) fertile - they can reproduce
Reproductive isolation
mechanisms that prevent gene flow between members of a different species\2 types
pre-zygotic or post zygotic
zygote formed by the fusion of to gametes
Pre-zygotic isolation (5 types)
1 - habitat isolation - ltwo species live in a different environment and never meet
2 - Temporal isolation - related species breed at different times of the year so never cross breed
3 - Behavioural isolation - different mating activities, males of one species are not attractive to the other
4 - Mechanical isolation - a mechanical mechanism that prevents cross breeding (eg flowers whos pollinator species cannot fit into to the other one)
5 - Genetic isolation - gametes cannot unite
Post - zygotic isolation (2 types)
Hybrid inviability - gametes unite but viable offspring cannot form
Hybrid infertility - gametes unite but offspring is infertile (mule)
Why is pre-zygotic more common than post zygotic?
Mating is costly:
Finding and attracting a mate takes time and energy
Producing eggs and offspring also takes time and energy
Individuals who mate outside their species produce fewer offspring and their alleles will be less represented in the next generation
Thus selection should favour those individuals who are better at identifying and mating with conspecifics (members of their own species)