Responses to Environmental Challenges Flashcards
How can strong light damage leaves?
Produces free radicals and reactive oxygen species (ROS)
Hoe can leaves protect themselves against reactive oxygen species?
- Xanthophylls cycle (preventing formation of ROS)
- Enzymatic destruction of ROS
- Rapid destruction and repair of D1 protein in PSII
What are photo-inhibition and photo-oxidation?
Photo-inhibition = damage due to strong light which is reversible via repair mechanisms
Photo-oxidaiton= irreversible damage
What are the relative capacities for damage repair in shade and sun leaves and why?
Shade leaves have a limited damage repair capacity and are more easily photo-oxidised
Sun leaves have a higher repair capacity (unless stressed by low temperatures) and do not usually become photo oxidised
Describe the xanthophyll cycle.
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How do chloroplast movements limit excess light absorption?
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What is dynamic photo-inhibition?
A method of temporarily diverting excess light absorption to heat whilst maintaining maximum photosynthetic rate
What are c3 and c4 plants
c3- cool season plants
c4- warm season plants
What is quantum yield? What is it dependent on and why?
Moles of co2 fixed/ moles of photons absorbed
Dependent temperature in c3 plants but nearly independent temperature in c4 plants, due to photosynthesis
What are the differences of photosynthetic capacities in c3 and c4 plants at different attitudes due to?
Due to reduced quantum yield and increased photorespiration
At which altitudes and attitudes does UV radiation decrease?
Higher altitudes and lower attitudes
How is UV radiation damaging to plants?
High quantum every damages. Long waveUV-A is photo oxidative. UV-B is photo-oxidative as well as causing photo lesions in biomembranes
Molecularly: UV breaks down disulphide bridges and dimerises thymine groups in DNA
What is dynamic photo inhibition?
The changes light response at different levels of light where a moderate level of excess light decreases the quantum efficiency without reducing the maximum photosynthetic rate
Describe dynamic photo inhibition and chronic photo inhibition compared to optimal photosynthesis
Moderate levels of excess light decreases quantum efficiency (reduced slope of the curve) without reducing maximum photosynthetic rate.
Whereas high levels of excess light
decreases quantum efficiency
and maximum photosynthetic
rate (chloroplast damage).
Give 4 ways in which leaves undergo photo protection as light flux increases.
- Decreased absorption
- Thermal energy dissipation
- Removal of ROS
- Inactivation/turnover of PSII (photosystem II)
What is the effect of co2 concentration leaf temperature?
Increased CO2 concentrations causes stomatal aperture to be smaller and therefore leaf temperature is higher as there is less transpirational cooling.
Describe the multilevel process of protection and repair of photo damage.
1) Suppression mechanisms e.g. using heat dissipation using the xanthophyll cycle
2) Detoxification- scavenging of ROS
3) repair mechanisms- synthesis of D1 protein
Describe the quenching process within photosystem II Why does this occur?
Violaxanthin is converted to an intermediate, antheraxanthin which is then converted into zeaxanthin (the fully quenched state)
This is done to use up excess energy that cannot be lost as heat or used in photosynthesis as converting between these forms uses energy
Describe the levels of zeaxanthin in evergreen leaves in winter compared to in summer.
Winter- high levels of Z maintained throughout day as they don’t know when they’ll have a cold day with high light intensities
Summer- levels of z rise and peak during the day to give more protection at noon