radiations and extinctions Flashcards
taxa
- heirarchial divisions of species from Kingdoms to subspecies
- group of organisms at any heirarchial rank
- e.g. family or genus
are smort or long-lived taxa more abundant?
short lived taxa are more abundant
what is the average age of species?
1-4 million years
what are background extinction rates?
number of species that would go extinct over a period of time based on non-anthropogenic (non-human) factors
- currently extinction rates are accelerating
- account for 95% of all extinctions
why do most organism go extinct?
- biological, climatic or physical changes to the environment
- multigenerational loss of reproductive fittness
- habitat attenuation and dissolution
- interbreeding (e.g. neanderthals interbreeding with homo sapiens caused their extinction)
- inability to compete with and displace the resident population
what is reproductive fitness?
how many offspring can be produced and their ability to reproduce
mass extinctions
- 5% of all extinctions
- when species vanish faster than they are replaced
- do not tend to be caused by cataclysmic events (e.g. earthquake)
- stochastic = population cannot recover once zero
- rapid = tend to occur within the lifespan of species
- higher survival chances when the geographical distribution is wide
what is a radiation?
- sudden loss in organism diversity leads to periods of expansion and diverification -> fill in vacant niches
- does not always occur due to an extinction
evolutionary radiation
- increase in taxonomic diversity caused by elevated rates of speciation
what can cause a radiation?
- extinction
- major transitions
- ecological specialisation
- major innovations
3 steps of ecological theory of adaptive radiation
- rapid diverisfication or organisms -> to exploit available ecological niches in the district (mutations accumulate)
- competition between different forms exploiting ecological niches
- specialization results from trade-offs & means that intermediate phenotypes might be selected against
what can cause adaptive radiations?
- ecological release = population increases when species are freed from limiting environmental factors
- ecological opportunity = environmnetal conditions allow niche availability, new ecosystems emerge
- key innovations / major transition
what is adaptive radiation?
diversification of a group of organisms where they fill different ecological niches
entropy
- measure of how energy is distributed in a system
- increased entropy = increased disorder
2nd law of thermodynamics
- entropy must always increase
- need to “cheat” this to be able to do things such as metabolism
local negative entropy
- increasing order
- achieved as long as the net entropy of a system is increased
- mini gradual dissipating systems have negative entropy -> osccilate between products and reactants
dissipation
energy not transferred to useful energy stores
energy wasted / lost to surroundings
when did only unicellular life exist?
3.8 billion - 900 million years ago
when did multicellular life forms emerge?
600 million years ago
how did multicellular organisms emerge?
- from unicellular ancestors
- under well defined environmental conditions
key specialisations of multicellular organisms
- cell-cell adhesion
- cellular specialisations
- germ-soma seperation
- alterations of life cycle via unicellular intermediates
selected synapamoprhies in animals
- diploid
- multicellular
- conserved genes for body plans
- posses true epithelia
- develop from a blastula
- eggs develop from 1 of 4 daughter cells in meoisis 2
- aerobic
- non-photosynthetic
- heterotrophic
what are synapamorphies?
characteristics present in an ancestor and is shared among the evolutionary descendants
what are the 2 layers of epithelial tissue found in all animals?
- endoderm
- ectoderm
endoderm
digestive system and organs like liver, lungs…
develops into gastrodermis (lining of gut cavity)
ectoderm
CNS and skin
develops into epidermis (outer layer od body wall)
what do the ectoderm and endoderm form?
A developing blastula
what is a blastula?
the early stage of an animal embryo
mesoderm
- 3rd layer of tissue only found in “higher” animals
- gives rise to muscles and organs between digestive tract and skin, supportive and contractile cells and blood cells
- found in triploblast organisms -> most triploblasts posses organ systems (e.g. digestive system, nervous system…)
protostomes
develop mouth first
deuterostomes
develop anus first
what are HOX genes?
- group of related and very conserved genes in animals
- transcription factors that control the body plan along the head-tail axis
- promote the transcription of certain genes
- vary due to genetic reshuffling
- abundant in different parts of the body e.g define if tail or no tail
what is the holobiant concept?
every animal is in a complex community
- bacteria cells outnumber host 10:1
- bacterial genome is larger -> more space for adaptations
body plans
group of characteristics shared by a group of phylogenetically related animals at some time
adaptive radiation in dogs
- dogs share ancestry with wolves and other canids
- dogs are derived from grey wolves only
- humans “selected” dogs that do well
canids
dog-like carnivores
not adaptive radiation in dogs example
- reproductive seperation and limited effective population sizes of established breeds
- opportunity for divergence by genetic drift
adaptive radiation in dogs example
- breeds under selection for morphological and behavioural traits
- selection due to humans
- breeds that do well are favoured
what forms the spectrum of carnial variation?
the combindation of adaptive and not adaptive radiation
- determines things like the bite force or breathing
modularity
- independence among groups of phenotypic traits
- if one changes, others are affected
- can help an organism avoid tradeoffs and likely alter the rate & direction of evolution
- may predispose dogs for rapid jaw length evolution
dog skull variation
- dogs have more diverse jaws and skulls than wolves do
- jaw comprises a module seperate from the cranium -> changes in jaw can be done without affecting the cranium
- jaw shape can change quite rapidly -> modularity -> due to very few genetic changes (which wont affect the whole skull)
dolichocephalic
long-faced breeds
brachycephalic
short-faced breeds
genetic basis for dog skull variation
- GWAS = at least 5 genetic regions / loci responsible for the cranioskeletal differences between dolichocephalic and brachycephalic breeds
- BMP3 mutation fixed among extreme brachycephalic breeds (pugs)
BMP (bone morphogenetic protein)
role in craniofacial development
central dogma of molecular biology
replication -> DNA -> transcription -> RNA -> translation -> proteins
morpholinos
molecules used & designed to block the activity of certain genes
what are species?
- DARWIN=set of individuals closely related resembling each other
- group of organisms with individuals capable of exchanging genes / interbreeding
- a principle taxonomic unit
- when at a certain point genes will no longer flow between two groups
what does it mean if an offspring is born highly altrical?
it is underdeveloped
will develop in the marsupium