Lecture Seven; Auditory organ Flashcards
What are the four parts of somatosensation?
Touch (cutaneaous senses)
Pain/temperature (nocicepsis
Position own bodyparts (propriocepsis
Organes (interocepsis
vestibular sense:
balance
The difference between eye and ear?
Ear is an analytic organ the eye is a synthetic organ
The ear is an analytic organ
(we perceive all individual frequencies)
The eye is a synthetic organ
(in a mix of light frequencies we perceive only a single color)
which cranial nerve is associated with hearing?
the eighth
what is analog to the brightness, hue (color), saturation the auditory organ?
saturation equals complexity (timbre), hue(wavelength) equals frequency (pitch) and brightness equals amplitude (loudness)
What is the tympanic membrane?
(eardrum),
which vibrates with the sound.
How is the middle ear structured?
The middle ear consists of a small hollow
region behind the tympanic membrane. It contains the
bones of the middle ear, called the ossicles, which are set
into vibration by the tympanic membrane. The malleus
(hammer) connects with the tympanic membrane and transmits
vibrations via the incus (anvil) and stapes (stirrup) to
the cochlea, the structure that contains the receptors. The
bottom of the stapes presses against the membrane behind
the oval window, the opening in the bone surrounding the
cochlea
Receptors in cochlea
Sound detection by bending inner hair cells in organ of corti
(outer hair cells are for amplification)
How are lower frequencies encoded vs. higher frequencies?
Detection high and middle high tones by “place coding “; low tones by “rate coding”
Rate coding:
tip vibrates at rate of low frequencies
Coding: intensity
High and middle high sound frequencies by neuron firing frequency (rate law)
But, not possible for low frequency sounds! For these the sound frequency itself is coded by the neuron firing rate.
Intensity of low frequency sounds is coded by the number of activated haircells
physical describtion of pitch and loudness
frequency and intensity/amplitude
Intensity of low frequency sounds is coded by …?
by the number of activated haircells
Coding location
Left-right by differences in arrival time in both ears
from right: right eardrum pulled in, left eardrum pulled out
from left: right eardrum pulled in, left eeardrum pulled out
from front or behin, botheardrums the same
why can we distinct from front and behind sound?
High-low (front-behind) changes in frequency-content induced by shape of ear shell
Higher order processing
In secondary auditory association cortices
- belt region (first auditory association cortex)
- parabelt region (second auditory association cortex)
- prmary auditory cortex
Information is combined for a complete percept
1. Where: Auditory Barking dog right (in front or behind?) Visual Nothing in front so behind
2. What: Auditory Rhustling in bushes (bird or mouse?) Visual Blue-yellow so bird
Perception of Pitch and Loudness High-frequency sounds and Moderate-frequency sounds
Perception of Pitch (frequency): Place coding; firing by hair cells at location of basilar membrane
that is active
Perception of Loudness (Amplitude)Determined by rate of action potentials from hair cells
Low-frequency sounds
Perception of Pitch :Rate coding; hair cells at apical end of basilar membrane
fire in synchrony with frequency of sound wave
Perception of Loudness: Determined by number of active hair cells
the four types of encapsulated somatosensory receptors
Merkel’s disks, Ruffini corpuscles, Meissner’s corpuscles,
and Pacinian corpuscles).
Merkel’s disks
Size and Nature of Receptive Field: Small, sharp borders
Location: Hairy and glabrous skin
Function: Detection of form and roughness, especially by fingertips
Ruffini corpuscles
Size and Nature of
Receptive Field: Large, diffuse borders
Location: Hairy and glabrous skin
Function: Detection of static force against skin; skin stretching; proprioception
Meissner’s corpuscles Glabrous
Size and Nature of
Receptive Field: Small, sharp borders
Location: glabrous skin
Function: Detection of edge contours; Braille-like stimuli, especially by fingertips
Pacinian corpuscles
Size and Nature of Receptive Field: Large, diffuse borders Location: Hairy and glabrous skin Function: Detection of vibration; information from end of elongated object being held, such as tool
Channels for positive ions (Na+) opened due to displacement rings: depolarisation
Open nerve endings
pain, temperature
Open nerve endings:
hair movement
Dermatome
A part of the skin subserved by one peripheral nerve
Shingles (Gordelroos):
viral infection of one nerve (dermatomes)
Projection to the cortex
Pain/temperature:
nociception
cogitive response:
Spino-thalamic tract (crosses immediately)
to Thalamus ventral posterior (passes Mediale lemniscus
)
and the to primary somtosensory cortex
Via various receptors on dendrites
Cognitive response
withdrawal reflex
Projection to the cortex
Touch:
Dorsal column then nuclei of the dorsal columns then also (as pain) Mediale lemniscus to Thalamus ventral posterior and then primary somatosensory cortex
tissue damage/inflammation
- macrophage:
- mast cells
- neutrophil granulocyte
through detection of signaling molecules
- Clearing damaged cells
- release of signaling chemicals
- Clearing damaged cells
Somatotopic map (place coding)
Somatosensory (and motor) homunculus