Lecture 18-27 Flashcards
ecosystem def
community of organisms and their physical environment
= all organisms in an area + physical envr + biotic/abiotic interactions
biome def
a grouping of ecosystems sharing a similar set of plant characteristics under a similar environmental regime
what are the two major sets of biomes?
terrestrial: primarily influenced by temperature, precipitation, seasonality
marine: primarily influence by water depth and proximity to land
What effects impact the general pattern of distribution of terrestrial biomes across latitudes?
temperature, precipitation, seasonality
distribution of biomes with increasing latitude is echoed with increasing elevation
- at the equator, it is hot and wet with low seasonality - we get tropical rainforests
- at 30 degrees, it is very warm and dry with moderate seasonality - we get desert
Warm air holds more moisture, therefore there are no biomes that have very low temperatures and high precipitation
What happens to NPP when temperatures and precipitation increase?
The higher the NPP, the more plants because there is an increase in photosynthesis (more water and sunlight)
Net primary productivity (NPP) differs between terrestrial biomes in a manner consisten with difference in temperature and precipitation
evaporation def
the movement of water directly to the air from the soil and water bodies
affected by heat, humidity and wind speed
transpiration
the movement of water from root systems through a plant, and exit into the air as water vapour
affected by plant type, soil type, weather conditions, and cultivation practices
evapotranspiration
transpiration + evaporation
on average, between 3/5 and 3/4 of land precipitation is returned to the atmosphere via evapotranspiration
how does deforestation lead to reduced evapotranspiration?
removal of vegetation (e.g. deforestation) decreases evaportranspiration (ET), and increases groundwater recharge (R), and river discharge (D)
(see picture on slide 17 of lect. 18)
How is NPP correlated with actual evapotranspiration (AET)
AET is affected by both temperature and precipitation
high AET=warm and wet
low AET=dry and cold, or both
Associate biome to NPP/AET
a) low NPP, low AET
b) medium NPP, medium AET
c) high NPP, high AET
a) desert, tundra
b) broadleaf forest, boreal/montane forest, dry tropical forest, grassland
c) wet tropics, wet temperate
Which type of forest accounts for about 1/3 of Earth’s terrestrial NPP?
tropical forests
give an approximate order of the biomes according to how much NPP they account for in increasing order
tundra, shrublands, boreal forests, deserts, crops, temperate grasslands, temperate forests, tropical grasslands/savannas, tropical forests
Which has a higher total NPP between tropical and temperate forests?
On a daily basis, the NPP per unit area is similar between tropical and temperate forests, this means that the difference in yearly NPP between the two biomes is primarily related to the length of the growing season.
therefore, tropical forests have a higher total NPP since their growing season is much longer than temperate forests
What plant characteristics define biomes?
Size, shape, foliage structure and chemistry of plants determine many ecosystem properties and the nature of the other biota
list the three general plant forms
grasses, shrubs, trees
disturbance def
events causing removal of biomass (e.g. herbivory, wind, frost, pathogens, erosion, fire)
competition def
ability to acquire resources compared to neighbours
stress def
any condition that restricts plant production (e.g., shortage of light, water, nutrients, or low temperatures)
Explain the relationship of disturbance, competition and stress with the three general plant forms
grasses: low competition, high disturbance, low stress
trees: high competition, low-medium disturbance, low stress
shrubs: low competition, low disturbance, high stress
(review this)
ruderals def
allocate resources mainly to seed production, often annuals or short-lived perennials
high growth rate, short-lived leaves, short statured plants
example: grasses
good competitors def
high growth rate, short leaf-life, low seed production, high allocation to leaf construction
example: trees
stress tolerators def
allocate resources to maintenance and defenses
often evergreen, long-lived leaves, low growth rate
example: shrubs
forests
trees are dominant or (co-dominant) plant type
two different types, based on longevity of leaf:
1) deciduous (1 growing season): winter-deciduous (subtropical and tropical, leaf shed on dry periods), drought avoidance
2) evergreen (>1 growing seasons): broadleaf-evergreen (tropic rainforest, no distinct growing season, year-round PS). needle-leaf evergreen (growing season is short or nutrient availability contrains PS and plant growth), drought tolerance
How much monthly precipitation provides sufficient moisture for plant growth?
about 20 mm of monthly precipitation for each 10 degrees celcius in temperature
Walter climate diagrams
permits ecologically meaningful comparisons of climates between localities
illustrates seasonal periods of water deficit and abundance
each climate zone has a typical seasonal patterns of T and P
Temperate seasonal forest
dominated by deciduous trees
soil is rich in organic material
challenges: high seasonality (hot summers, cold winters)
deciduous leaves change colour and fall during autumn, thick bark (protection), shade-tolerant understory, lots of leaf litter
boreal forest
dominated by coniferous forests
soil is acidic and mineral-poor
challenges: shorter growing season, long cold winters
conical shaped conifers, dark coloured needles (less surface area –> less evaporation), waxy coating (reduced evaporation), seeds in protective cones
tundra
soil extremely rich and organic
challenges: short growing season, permafrost, extreme cold, poor drainage, very windy
shallow root systems, low to the ground, dark colours, grow close together, lots of lichen and moss, small leaves, wax fuzzy coating, most are perennials
temperature grassland
seasonality: moderate (hot summers, cold winters)
soil is extremely rich and organic
challenges: frequent fires, droughts, windy, grazing
narrow leaves, soft stems, extensive root systems, leaves contain silica, grow from near their base, wind pollination
tropical rain forest
soil is low in organic content
challenges: bacteria and fungi, risk of flooding and erosion of soil and leaching of soil nutrients
colourful plants/flowers attract pollinators (since no wind)
drip trips and waxy surfaces
broadleaf evergreens
smooth thin bark
subtropical desert
precipitation very low <25cm/year
soil is mostly sand (90-95%); low N and organic material; high CaCO3 (calcium carbonate) and phosphate
challenges: windy (no cover), extreme heat and drought and large temperature shifts (cold at night)
waxy coating, thick juicy leaves
white hairs
bloom at night
expendable stem
spines insteawd of leaves
What are the world’s most impacted biomes?
tropical dry forests and temperate grasslands
(56% terrestrial surface (minus permanent snow and ice) has low human impact)
Profound principles behind marine biomes
Life is short
things sink
it gets dark down there
and cold
What influecnes primary productivity in the ocean?
Availability of nutrients (N,P, Fe, Si)
Amount of sunlight
Ocean stratification
Heat arrives at the ocean surface from above
90% of radiation entering ocean is absorbed in the top 100 m
Warm water is less dense than cold; warm water on top of cold is a stable configuration
Surface +/-200 m is well mixed and separated from deeper water
Surface currents and deep currents behave differentlt, although they are linked
Top layer floats on top and cold water below (warm water is less dense)
Ocean layers
top 150m: warm, nutrient-depleted surface water
150-250m: thermocline
bottom: cold, nutrient-rich deep water (because not much competition for the nutrients)
productivity in temperate oceans (according to seasons)
winter: short days, little light (no phtosynthesis), low productivity (missing warm temperature), not much thermocline (all cold in the winter)
spring: phytoplankton bloom, but quickly uses up the nutrients and then low productivity, solar radiation available for photosynthesis, not much thermocline so mixing occurs
summer: strong thermocline means that when nutrients are used up, there is a low productivity, strong thermocline keeping cold nutrient rich water below
fall: thermocline breaks down, bringing nutrients up from below, so we have a short-lived phytoplankton bloom, thermocline begins to disappear, secondary bloom
Comparing productivity of polar midlatitude, and tropical ocean regions
polar regions: extremely high rate of productivity during the summertime
tropical regions: steady, low rate of productivity year-round
middle latitude: large peak in productivity during the spring and a lesser peak in productivity during the fall
What are the marine biomes, and how are they distributed?
Light availability:
- euphotic: top layer of ocean, most sunlight
- photic
- aphotic: no more light
Water depths
- pelagic: open water, not on the bottom, surface all the way bottom excluding the floor
- benthic: floor of the ocean
- abyssal: on the bottom, but very deep
Distance from land:
- intertidal: mangroves
- neritic (continental shelf): coral reefs, kelp forests
- oceanic: all the rest of the ocean away from the continental shelf
Euphotic zone
topmost part of the ocean where light is the strongest
where nearly all of primary production from photosynthesis occurs
photic zone
upper part of the ocean where light penetrates
aphotic zone
Lower part of the ocean where very little or no light penetrates
No living plants
High pressure, low temperatures
Animals survive by eating detritus or other animals
Must adapt to living with no light
Is NPP high or low in open ocean
NPP is lower in the open ocean relative to the shore because nutrients are low
Where does photosynthesis peak in the ocean?
Approx 100m below the surface
Photosynthesis is inhibited at the very top since there is too much light
Lower light levels restrict photosynthesis down here
Benthic zone
Any sea bottom surface
Low oxygenation of water
Low temperatures
Animals here feed on detritus or other animals
Little or no plant life (depending on water depth)
Abyssal zone
Subdivision of benthic zone
Floor of the deepest parts of the ocean
Incredible water pressure
Absolutely no light
Very cold temperatures
Hard to survive
Pelagic zone
open ocean of any depth
Why is depth is a key factor in determining the distribution of the dominant marine biomes?
Depth determines:
- Light intensity (photic zone=high light; photosynthesis>respiration)
- Temperature
- O2 concentration (depends on temperature and balance between photosynthesis and respiration)
- Pressure
- Nutrient concentrations (atmospheric input at surface; nutrients consumed and tied up in organisms near the surface, released in aphotic zone by bacterial decomposition)
Organic matter in relation to depth
Organic matter is high near the surface, and is depleted and nutrient concentrations are enriched with increasing depth.
Most life is in shallow ocean where photosynthesis happens. When things die, they usually get consumed before sinking too low
Oxygen in relation to depth
oxygen is high at depth due to deep current sources and low temperature (conveyor belt)
How are nutrients brought to photic zones?
Upwelling along continental margins brings nutrient rich waters into the photic zone; high nutrients, oxygen and light support high productivity
As you go deeper, what happens to:
a) light/temperature
b) salinity/density
c) nitrates/carbon
d) oxygen
a) decreases
b) increases
c) increases
d) decreases, and then increases
Why is distance from land a key factor in determining the distribution of the dominant marine biomes?
input of nutrients to photic zone
- nutrients in airborne dust
- nutrients from rivers and estuaries
- nutrients from coastal upwelling
Intertidal zone
Where land and ocean overlap
Abundant sunlight
Constant wave action supplies nutrients and oxygen
Food is abundant
Varied substrate provides hiding places and surfaces to cling to
Neritic zone
seaward from the low tide line, the continental shelf out to the shelf break
Well oxygenated water
Low water pressure
Stable temperature and salinity levels
Home to photosynthetic life
Oceanic zone
Beyond the continental shelf
Larger creatures
Life decreases with increasing depth
Widest array of life (because it is a very broad area)
Upwelling def
process in which deep, cold water rises toward the surface (drive up cold deep ocean water filled with nutrients)
Perks to intertidal zone
Abundant sunlight
Abundant nutrients and oxygen (waves, proximity to land)
Varied substrate for hiding places and surfaces to cling to
Challenges to intertidal zone
rapidly changing conditions - exposure to surface and sun varies, salinity
Abundant sunlight can lead to desiccation
Waves can carry you away
Competition for space, light, food
Exposed to predators when tide is out
Name some adaptions to life in intertidal zone
sticky podia (seastars)
Byssal threads (mussels)
close shells tightly (barnacles)
Holdfast root system - attaches to rocks and mussels (kelp)
CaCO3 structure, red colour
Cluster and secrete mucus (snails)
Neritic zone characteristics
Shallow, sunlit waters
Plentiful nutrients from land and upwelling
Most productive and economically significant parts of ocean
90% global catch of shellfish and finfish from here
Where is productivity the highest and lowest in marine environments?
highest: continental shelves
lowest: open ocean
Species def
a group of genetically similar organisms that can interbreed and produce fertile offspring
(def does not apply to asexual organisms)
species diversity
typically measured as species richness: # of species within a habitat
How does species richness change towards the tropics?
increases toward the tropics
1) tropics have greater land mass: more area, therefore more species (however, polar regions have fewer species than similar areas in lower latitudes)
2) harsher climates at the poles
3) tropical regions have more stable climates (but also applies to top of mountains)
4) species-energy hypothesis: greater exposure to solar energy –> greater productivity –> more species can be accomodated in food webs
5) higher speciation rates (driven by energy), greater accumulation of species over evolutionary time
What are the functional differences between terrestrial and marine biomes?
precipitation/temperature:
- key controlling factors in terrestrial biomes
- precipitation unimportant and temperature less important in marine biomes
Variation in seasons:
- high variation between seasons in terrestrial
- low variation between seaons in marine
Organism adaptions:
- terrestrial: organisms exposed to extremes of climate; adapted to moisture/temp. regimes
- organism adaptions similar in all marine biomes
materials:
- terrestrial: fall but largely remain in the ecosystem; recycled
- marine: sink=become inaccessible to organisms at a given depth; “biological pump”
nutrients:
- terrestrial: via soil and atmosphere
- mainly from atmosphere (precip. and dust); proximity to land is important
Effect of gravity on terrestrial vs. marine organisms
Land plants tend to have structures to live with gravity (structures that are harder to breakdown and decompose)
Gravity is not a factor when you are in the water, structure is squishy, as a result get eaten more easily and quicker
What does water availability depend on?
Residence time in the reservoir
Temporal availability
Spatial availability
Proximity to human need (availability vs. access)
Residence time def
length of time water remains in a reservoir differs
averages can be deceiving
- different reservoirs of the same type can have different residence times
How is freshwater classified based on residence time?
renewable:
- water in the reservoir is continuously recharged (<1 year)
- significant withdrawals do not typically cause depletion within the reservoir
- can be depleted if withdrawals are significant enough
non-renewable
- reservoirs which are not recharged on human time scales, or recharge so slowly that significant withdrawals will cause depletion within the reservoir
Temporal availability
not all freshwater is equally accessible year round
- rainfall patterns vary seasonally
major changes in seasonal rainfall patterns
- rainy season and dry season
Spatial availability
rainfall patterns differ globally
majority of precipitation in the equatorial region (30 degrees north and south of the equator)
frequency of terrestrial reservoirs differs (both for green and blue water)
local proximity (location of freshwater resources must be within reach of the human activities requiring the water resources
available freshwater resources and population is unevenly distributed in Canada (issues with water security)
Available water
how much water is available for potential usage
accessible water
how much available water we can actually access for use
- spatial and temporal location
- economic access
- culturally and politically acceptable access