Lecture 17 Flashcards
Protochordata, Craniata, Gnathostomata
All are within Phylum Chordata
- Protochordata
- Do not have a true brain, skull, or vertebral column
- Craniata (i.e. Vertebrata)
- Have a skull (cranium) and/or vertebral column
- Agnatha (Hagfish and Lampreys) only have a skull (no vertebrae)
- Gnathostomata
- Craniates with jaws
- Includes all of Craniata except for Agnatha
Phylogeny of fish!
fish itself is any non tetrapod vertebrate
Agnatha
Osteichthyes* (Bony Fishes)
Chondrichthyes
General characteristics to distinguish what is a fish (5)
- Form
- Locomotion in water
- Neutral buoyancy and the swim bladder
- Respiration
- Osmotic regulation
Form – types of scales for fish
- Placoid scales
- Ganoid scales
- Cycloid and Ctenoid scales
Placoid scales
- Small, conical, toothlike structures
- Typical of chondrichthyes
- Modified to teeth in sharks
Ganoid scales
- Diamond shaped
* Early bony fishes and living gars
Cycloid and Ctenoid scales
- Arranged in overlapping rows
* Typical of teleost fish
Locomotion in Water for fish
Movement achieved through undulation of the posterior end
• Generates thrust (forward motion) and lateral force (sideways motion)
• The lateral force causes fish head to “yaw”, or deviate in the same direction as the tail
• The “yaw” occurs more in more flexible fish
• A less flexible body plan is conducive to speed!!!!!!!!!!!!!!!
Buoyancy in fish
Swim bladder = gas-filled organ
Volume adjusted for neutral buoyancy
present in most pelagic (open sea) bony fishes
Not in very deep fish, and most bottom-dwellers
Why don’t Chondricthyes (e.g. sharks) have no swim bladder
- Asymmetrical tail provides lift,
* Large livers with squalene (particularly buoyant lipid)
Respiration for fish
Most fish use gills
• Breathe dissolved oxygen (oxygen that is in the water)
• Gills are located in the pharyngeal cavity
• Gills are covered with an operculum in bony fishes
• Increases efficiency of respiration
• Not present in sharks and rays
• Some fish also have lungs ,Capable of breathing air
Gills in fish
- Gill composed of thin filaments covered with an epidermal membrane
- The membrane is folded repeatedly into plate-like lamellae
- Enormous surface area
- Lamellae contain main blood capillaries
- Water is pumped continuously in the mouth, over the gills and out through the gill slits
- Gas exchange occurs across thin walls of blood capillaries
Osmotic Regulation in fish
- Maintenance of balance of fluids
- Freshwater and Marine fish have opposite challenges
- Freshwater fish are hyperosmotic regulators
- Marine fish are hypoosmotic regulators
Hyperosmotic Regulators
Freshwater fish
• Greater salt concentration in fish than in surrounding water
• Scales and mucous protect the fish, but water can enter across membranes (e.g. gills)
• Water pumped out by kidneys
• Salt-absorbing cells in the gill move salt from water to blood
Hypoosmotic Regulators
- Marine fish
- Contain a smaller concentration of salt than surrounding water
- Salt-secretory cells in the gill move salt out of body
- Salt is voided with feces or excreted by the kidney