Final Flashcards
What is a community?
all populations of species living together in a
particular area
Challenges of measuring an ecological
community:
- Difficult to count ALL species present
-Therefore, often specify - all plant species, frugivore community - Community boundaries often gradual
- Species can move in and out of communities
What do we mean by Community structure?
1) which species are in the community
2) what is relative abundance of each species
3) what are the relationships among the species
Ecotone
-changes in the environmental conditions
-ex: soil type, geology, water
- aspect, disturbance (grazing, plowing)
a boundary created by sharp changes in environmental conditions over a relatively short distance, accompanied by a major change in the composition of species
Detecting an ecotone:
run a transect from non-serpentine to
serpentine
- note which species are present at each point
along the transect - ecotone – where we see a shift from one set
of species to another
What is serpentine soils?
low nutrients, high in metals
interdependent communities
communities in which species depend on each other to exist
independent communities
communities in which species do not depend on each other to exist
-species just happen to live together cuz they have similar requirments/adaptions
What is species richness?
the number of species in a community
Why is species richness often higher at an ecotone than not at an ecotone?
-generalists can live there and some of the species from each of the 2 communties xan live there
- overlap between communties
Community structure:
- Which species are in the community?
- What is the relative abundance of each
species? - What are the relationships among species?
abundance:
the number of individuals
relative abundance:
the proportion of all individuals represented by each species
What is species evenness?
-evenness is highest when all species have equal abundance
a comparison of the relative abundance of each
species in a community
Rank-abundance curves
plot the relative abundance of each species in a
community in order from most to least abundant
steeper = less even
never a complete even species
log-normal distribution
a normal, or bell-shaped, distribution that uses a log scale on the x-axis
Diversity indices:
a way to compare the diversity of communities that takes into account species richness AND evenness
Simpson’s index:
a measurment of species diversity, given by formula on page
Shannon’s index:
a measurment of speices diversity given by formula on page
intermediate disturbance hypo
found that hump-shaped relationships found in <20%
the hypo that more species are present in a community that occasionally experiences disturbances than in a community that experiences frequent or rare disturbances
(1) Productivity (resources)
species richness can be affected by the the amount of resoucres available (soil nutrients)
- productivity is measured by biomass = more resources
- relationships found in nature between productivity and species richness: hump-shaped is most common
Ecologists have found that-
as more and more resources are added, the number of plant species goes down
What causes more and more resources that are added, the number of plant species goes down
at high levels of resources, it allows a few dominant competitors to outcompete the other species
whether the relationship between productivity and species richness looks positive, negative, or hump-shaped can depend on -
the RANGE of productivity in the experiment
Habitat diversity
- communities with a greater diversity of habitats
-habitats tend to have more species
- more niches available
Keystone species
- the presence of a keystone species can lead to greater species richness (even if that species is not very abundant)
Disturbance
Can also affect species richness in a community
ex: hurricanes, wildfires, logging, ploughing
ecologists agree that disturbance (type, severity, frequency) affects community structure
- but the IDH is controversial
What local factors can affect community species richness?
(1) The amount of resources (productivity)
(2) Diversity of the habitat
(3) Presence of keystone species
(4) Disturbance intensity or frequency
food chain
a linear representaion of how different species in a community feed on each other
food web
a complex and realistic representaion of how psecies feed on each other in a community
-who eats whom?
Why do ecologists want to know who eats who?
-understand feeding relationships
-how one species change in abundance can affect another
-weather a species can survive in an area
-what would happen if one species goes extinct
Trophic levels:
a level in a food chain or food web of an ecosysytem
- all species in a trophic level get their energy
in a similar way
When discussing trophic levels, a guild is:
a group of species that feed on similar items.
Do species in the same guild have to be closely related?
no,
-ex, ants and rodents
primary consumer
a species that eats producers
secondary consumer
species that eats primary consumers
tertiary consumers
a species that eats secondary consumers
omnivore
a species that feeds at several trophic levels
Types of indirect effects
Density-mediated
Trait-mediated
Density-mediated
- caused by changes in the DENSITY of intermediate species
-fish eat dragonfly larve = less dragonfly density= less predation on pollenators
Trait-mediated
- caused by changes in the TRAITS of the intermediate species
-mediated
-behaviour
indirect effects
an interaction between two species that involves one or more intermediate species
direct effect
an interaction between two species that does not involve other species
trophic cascade
indirect effects in a community that are initaiated by a predator
density-mediated indirect effect
an indirect effect caused by changes in the density of an intermediate species
trait mediated indirect effect
an indirect effect caused by changed in the traits of an intermediate species
community stability
the ability of a community to maintain a particular structure
community resitance
the amount that a cmmunity changes when acted upon by some disturbance, such as the addition or removal of a species
community resilience
the time it takes after a disturbance for a community to return to its original state
alternative stable state
when a communtity is disturbed so much that the species composition and relative abdundance of populations in the community change, and the new community structure is resistant to further change
bottom up effect
happens at the bottom and affects all
top down effect
or “tropic cascade”
what regulates populations?
both top down and bottom up together
(1) the world is green (plants are doing well!), therefore, something must be controlling the herbivore populations
(2) that must be the predators
(3) therefore predators are the most important (top-down control)
Succession:
the process by which the species composition of a community changes over time
Seral stage:
each state of community change during the process of succession
e.g. early seral, late seral, climax community
Climax community:
the final stage in succession
does not mean permanent or totally stable
Pioneer species:
the first species to arrive at a site
-get succession going (partly started)
how can we study succession?
1) direct observation
2) indirect observation
chronosequence (indirect)
a series of sites that differ in age since abandonment or disturbance, but otherwise occur on similar soil types and environmental conditions
land rising up gradually
-farther from water= older communities
direct observation
-keep returning to the same site overtime and record which species are there
ASSUMPTIONS for Chronosequence
1) every site foes through the same stages
2) the sites vary in AGE only
ex- same climate, same rock underneath
Indirect methods
- Chronosequence (space for time)
- Paleoecological methods
Paleoecological methods
can use instead of chronosequence
Paleoecology is the study of the relationship between extinct organisms and their environments.
-pollen in lake sediments
-tree rings
Observe it directly over time pros and cons
PRO: no assumptions of space-for-time
CONS: 1) have to wait a long time (especially long species like trees)
2) maybe have only 1 example (n=1)
Indirect methods pros and cons
PRO: takes less time!
CONS: 1) have to make assumptions
2) paleoecological proxies have variable resolutions
- ex, cant differentiate to species, maybe only to genus using pollen for trees
Why might direct observation of succession be easier in intertidal communities than in terrestrial communities?
Organisms in an intertidal area have shorter generation times than organisms in terrestrial communities.
Types of succession in terrestrial
environments:
Primary succession
Secondary succession
Primary succession
“start from scratch”
- habitats start with NO plants and NO organic soils
sand dunes, lava, bare rock, volcanic ash
Secondary succession
habitat has been disturbed, no plants but DO have organic soils and could have seeds or roots
plowed fields, forest fires
Species traits: for early seral
seeds = many
seed size = small
dispersal mode = wind/ stuck to animals
seed viability = long
root:shoot ratio = low
Growth rate = fast
Size = small
Shade tolerance = low
Species traits: for late seral
seeds = few
seed size = large
dispersal mode = gravity/ eaten by animals
seed viability = short
root:shoot ratio = high (many roots)
Growth rate = slow
Size = large
Shade tolerance = high
What causes succession?
Why does the composition of the community shift over time?
-condtions are changing (light, soil)
-caused by other species
Facilitation:
Inhibition:
Facilitation:
the presence of one species INCREASES the probability that a second species can become established
-ex, alder trees, n-fixing creats greater n availabliity in soil, which allows spruce to colonize
Inhibition:
the presence of one species DECREASES the probability that a second species can become established
ex- maple and beech trees casting deep shade prevents early seral from growing there
What is NOT an example of inhibition?
A species acts as a nurse plant and increases water available to other species.
(its helping so no)
How can you determine whether a particular species is FACILITATING or INHIBITING another species? (or, neither?)
HYPOTHESIS: acorn barnacles are facilitating macroalgae
if the hypothesis is true, then areas with acorn barnacles removed will have significantly lower density of algae than the control after 2 years
“Climax” communities are not unchanging
-small disturbances can create shifts in species composition
- e.g. old tree dies, falls = GAP
- gap can be colonized by early-seral species
- community overall is in late-seral stage, but has some early-seral species due to the gap
what is a GAP
pockets of early succession
priority effect
when the arrival of one species at a site affects the subsequent colonization of other species
transient climax community
a climax community that is not persistent
fire-maintained climax community
a successional stage that persists as the final seral stage due to periduic fires
grazer-maintained climax community
when a successional stage persists as the final seral stage due to intense grazing
Primary productivity:
The rate at which solar or chemical energy is captured and converted into chemical bonds by photosynthesis or chemosynthesis
Standing crop:
the biomass of producers present in an ecosystem in a given area and at a particular moment in time