Final Flashcards
- All early hominin fossils, older than 2 mya are found.. 2 main regions
in Africa.
Parts of East and South Africa.
All early hominin fossils
*why Africa?
- Likely larger populations lived in this area than other areas.
- Geological features may help them preserve better.
- Caves in South Africa:
- Taung- Taung child cranium 1925
- Sterkfontein
- Fromfraai
- Swartkrans
- Malapa- recently, found in 2008.
- These have been made UNEXCO Heritage site called Cradle of Humankind
Dolinas
- Vertically oriented caves
- Formed by dissolving limestone, eroded bedrock.
- Over millions of years, these grows large and deep into large cavities with a small opening at the top.
- Stratification happens in the cave.
- Some used by hyenas.
- Some hominin groups used Dolinas for living.
- Breccia-
new rock that forms over time.
* 10-40 meters deep.
- Sterkfontein
20 m of breccia deposits. Date to 3.5 million years ago at the bottom and 200 000 years ago at the top.
* Over 700 hominin bones and fragments have been found here.
* Almost complete skeleton was found here, same as Taung child.
* Need to use very small tools as its wedged in there.
Australopithecus
- Malapa
- Mostly complete hominin skull found date to 2 million years ago.
- .
Australopithecus sediba
- More sites in East Africa
More sites in East Africa than south Africa
* Tanzania- Olduvai Gorge, Laetoli
* Kenya- West Turkana, Koobi Fora
* Ethiopia- Hadar and Kada Gona.
* Not individual sites, but large areas on the landscape where fossils are found.
- Olduvai Gorge-
- 2 million years ago it was a lake, and hominins and animals lives along the shore
- Found stone tools and bones, phytoliths- plant matter from fossilized palm trees.
- Used to have lake but now dry and lots of erosion- so that means fossils are able to be found.
- Lousi and Mary Leaky started finding fossils in 1959
- Laetoli
- Famous for Mary Leakey’s discovery of hominin footprints in volcanic ash from 3.5 million years ago.
- Over 70 footprints from 5 individuals, give info about walking gait at that time. Many animal footprints found as well
Australopithecus
- Koobi Fora
- Animal and hominin footprints
- 1.5 million years old.
- Genus homo fossils found here- most complete ones
- West Turkana
- Old river deposits to 4 million years ago.
- Stone tools sating to 3.3 mya? Possibly- would be the oldest artifacts ever found.
- Afar Depression- Ethiopia.
- Used to be thick forest and wet marshy areas
- Hadar
- Rich in hominin remains, date from 4.5 to 3 million years ago
- Ongoing erosion changes the landscape and makes it so that fossils can be found even if the area was already searched.
Lucy found here
- Kada Gona
- Previously oldest stone tools 2.6 million years ago
- What is a Continental Rift?
- A linear zone along which continental lithosphere stretches (and thinned) and pulls apart. Its creation may mark the beginning of a new ocean basin
- Tensional forces and buoyant uplifting of the heated lithosphere cause the upper crust to be broken along normal faults, while the lower crust deforms by ductile stretching.
- As the crust is pulled apart, large slabs of rock sink, generating a rift valley
- Further spreading generates a narrow sea
- Eventually an expansive ocean basin and ridge system are created.
- The East African Rift
is a continental rift that extends through eastern Africa for approximately 2,000 miles. It is made of several interconnected rift alleys that split into eastern and western sections around Lake Victoria. Eventually, a section of Africa will be divided by ocean.
- What causes Rifting?
- Mantle Convection
- Plumes
- Slab Pull and Ridge Push (subduction)
- Gravitational Collapse
- Started 24 MYA
- Rift in Malawi opening at less than 3 mm per year.
- What is the geological effects of the African Rift?
- Creation of a ocean basin.
- The opening of a new ocean basin begins with the formation of a continental rift. When rifting continues, the rift system evolves into a young, narrow ocean basin.
- Earthquakes.
- Any movement within a fault system can cause strain that builds up and result in devastating earthquakes
- Rift Volcanoes.
- Rift volcanoes form when magma rises into the gap between diverging plates.
- Fossil Hominid Sites of South Africa The Taung Skull Fossil Site
- :
- 1924 the celebrated Taung Skull – a specimen of the species Australopithecus africanus – was found
Makapan Valley, also in the site, features in its many archaeological caves traces of human occupation and evolution dating back some 3.3 million years
Fossils found there have enabled the identification of several specimens of early hominids, more particularly of Paranthropus, dating back between 4.5 million and 2.5 million years, as well as evidence of the domestication of fire 1.8 million to 1 million years ago
The Earliest Fossil Hominins
Extinct or fossil species that are linked to us.
A couple of important terms:
Plesiomorphic Trait A primitive trait inherited by a species from their ancestor.
prehensile feet in living apes
Apomorphic Traits New or ‘derived’ trait. It first appears in the species in question. Not inherited from an ancestor. E.g. Early Miocene apes had no tail. Likely as they were in trees less so didn’t need it. They did evolve from monkeys with tails tho.
no tail in primitive apes- no tail in humans.
non-prehensile feet in later hominins
Plesiomorphic Trait
A primitive trait inherited by a species from their ancestor.
Apomorphic Traits
New or ‘derived’ trait. It first appears in the species in question. Not inherited from an ancestor. E.g. Early Miocene apes had no tail. Likely as they were in trees less so didn’t need it. They did evolve from monkeys with tails tho.
no tail in primitive apes- no tail in humans.
non-prehensile feet in later hominins
When researchers dig up a new fossil …
They need to decide if it is … ??
Where does it fit into our evolutionary tree?
* just another example of something we’ve already found before?
* or a new species?
* or even a new genus?
* Need to study the features of the fossil.
Evolutionary Systematics
process of constructing the evolutionary relationship of the fossuls.
Phylogeny
refers to the actual evolutionary relationships between different organisms.
taxonomy
the process of classifying organisms, based on available data, about their phylogenetic relationships
With living organisms (and a few fossil species) we can use DNA to get a more accurate idea of actual evolutionary relationships (as we saw with the living primate taxonomy) gives us almost 100% confidence. But for super old ancestors, we do not have DNA.
This is mainly based on how different or similar two fossils are in their morphology – e.g., what plesiomorphies do they share and what apomorphies distinguish them? * different groups in the taxonomic system (different species or different genera) are referred to as different taxa (singular = taxon)
Pliocene
Genus Homo (hablis) The Pliocene is a geological epoch that occurred from approximately 5.33 million to 2.58 million years ago, preceding the Pleistocene epoch.
Pleistocene
Genus homo turn into homo erectus, neandertals
2.6 MYA to 12k years ago
Holocene
started 12000 years ago- switch from hunting and gathering to farming etc.
writing starts?
Last half of Holocene
The last 7 million years … Important geologic time periods
The last 7 million years … Important geologic time periods:
The last 7 million years of Earth’s history fall within the Quaternary Period, which is part of the Cenozoic Era. The Cenozoic Era began around 66 million years ago and continues to the present day. The Quaternary Period is characterized by the presence of repeated glaciations, and it is divided into two epochs: the Pleistocene and the Holocene.
Pleistocene Epoch (2.58 million years ago - 11,700 years ago):
The Pleistocene is often referred to as the Ice Age. During this epoch, Earth experienced multiple glacial and interglacial cycles, with ice sheets advancing and retreating. These climatic fluctuations had a significant impact on the landscape and the evolution of life.
Holocene Epoch (11,700 years ago - present):
The Holocene represents the most recent geological epoch and is characterized by a relatively stable and warmer climate compared to the Pleistocene. It encompasses the time since the last major ice age, and it includes the development of modern human civilizations. The Holocene continues into the present day.
These geological time periods are part of a hierarchical system used by geologists to categorize Earth’s history based on significant events and changes. The Quaternary Period, with its Pleistocene and Holocene epochs, captures the environmental and biological developments of the last 7 million years.
Hominid’ vs ‘Hominin’
Hominid:
Historically, the term “hominid” was used to refer to all members of the biological family Hominidae. This included not only humans but also great apes such as chimpanzees, bonobos, gorillas, and orangutans. In this broader sense, hominids encompassed all the great apes and their ancestors.
Hominin:
The term “hominin” is a more specific and modern classification. It is used to describe the tribe Hominini, which includes modern humans (Homo sapiens) and our closest ancestors after the split from the common ancestor with chimpanzees. In other words, hominins include all species more closely related to humans than to chimpanzees.
The key distinction is that “hominid” includes a broader group that comprises both humans and great apes, while “hominin” is a more narrow classification specifically referring to humans and their closest extinct relatives.
- Quaternary period
- Quaternary period is what has the Pleistocene and Holocene epochs.
Hominini
the tribe level which we separated from apes.
* Chimps and bonobs and us come from Panini. Use chimp as an analog for what early homnin used to look like.
Chimpanzee post-crania:
Unlike us
* long arms
* short legs
* long, narrow pelvis
* legs widely spaced
* long narrow fingers/toes
* prehensile hands and feet
* narrow from side to side, deep from front t back rib cage
Chimpanzee cranium: * brain size: 300-400 cc
* robust browridge
* alveolar prognathism- lower face protrudes.
* canine pillars- bony structure in bower face- provides support for powerful bite.
* u-shaped tooth row
* small molars & premolars – only slightly larger than ours.
* large incisors
* very large canines w honing complex (when close mouth upper caninie is sharpened against the premolar)- especially upper and in males.
Last 8 million years- most important- we split from apes.
Sahelanthropus tchadensis
Sahelanthropus tchadensis is an extinct hominid species that lived approximately 7 million years ago during the late Miocene epoch. The fossilized remains of Sahelanthropus were discovered in Chad, Central Africa, at a site known as Toros-Menalla.
7 mya “Toumai” means hope of life. South sarahan desert in Chad. – 7 MYA- around the time of homonin chimp split. Has ape like face and small brain- comparable to chimps. Could be a chimp ancestor. Do not know how bipedal it was. Nuchal line on back of skull- where neck muscles attach to cranium. This muscle is larger and attaches higher on skull if they are quadrupeds. In humans it’s a little lower. But Toumai may be bipedal based on the location of the nuchal muscle.
Foramen magnum
- related to posture- humand in centre of base of skull- because we stand upright. Quadruped has this at back of skull. Chimp is halfway between. Toumai is kinda in the middle of humans and chimps so not sure if bipedal.
Orrorin tugenensis
… an obligate biped? ≈ 6 mya East Africa A more habitual biped than chimps and bonobos ..Around the time of ape-human split.
Similar length to chimps- but was likely more bipedal than chimps based on bones in leg. Cant say for sure, need more fossils. Good candidate for being early hominin.
From 5 to 1 mya (million years ago):
another important hominid species in the study of human evolution. It lived approximately 6 million years ago during the late Miocene epoch. The fossilized remains of Orrorin were discovered in the Tugen Hills of Kenya, East Africa.
one of the oldest known hominids. It dates back to around 6 million years ago, placing it in the late Miocene epoch.
considered by some researchers to be a potential candidate for a common ancestor of humans and chimpanzees. Its bipedal adaptations and age make it an important species for understanding the early stages of hominid evolution.
4 genera and up to 20 species (we’ll just look at the major ones) our direct line
Ardipithecus: Ar. ramidus
Australopithecus: A Anamensis, a. afarensis. A. africanus, A sediba
Paranthropus: P boisei, P robustus, P aethiopicus
Homo: H. habilis, H ergaster, H. erectus.
Ardipithecus ramidus
4.5 mya. Discovered in 1992 in Ethiopia. Much larger than chimp cranium. Brain similar size to chimp 300-400 cubic cm. Large incisors, molars are rectangular- which is apelike. Reduced ale ho prognathism protruding face. More bipedal.
Strongly arboreal- prehensile feet.
ATM oldest hominin ancestor
Reduced ‘prognathism’
Australopithecus anamensis
4.0 to 3.0 mya East African Species. First found my Maeve Leaky in 90s. no complete crania. Small brain 370 cc. Apelike cramium and dentition. Large caniniens, signifigane sexual dimorphism.
Lower leg bones- tibia, indicates bipedal and no prehensile feet.
Apomorphic features?
Feet that are no longer prehensile Lower leg (tibia) Apomorphic features? Fully obligate bipeds
Australopithecus afarensis- Australopithecus africanus
≈ 3.0 - 2.4 mya South likely our direct ancestor. East Africa
African Species
afarensis and
africanus- sputh African species.
Both had small bodies, dexual dimorphism- males are 30% larger than females. 110 cm vs 140 cm. likely lot so fo competition for females.
Both craniums are about 425 cc. not much bigger than chimp and less than 1/3 the size of our brains.
Canini diastema (diastema, or a gap, between their canines and adjacent teeth) disappears in all hominins after this.
Australopithecus sediba ?
2.0 mya South African Species (Malapa)- not called species till 2010. Malapa 1- young boy Malapa 2- adult female. 420 cc brain, 130 cm tall. Fossils spread out in the breccia- not sure if all the bones are just these 2 individuals. We are unsure of this. Could be a mish mosh of other species and that’s why they look new.
Australopithecus africanus
≈ 3.0 - 2.4 mya South African Species
afarensis and africanus
‘Dental Arcade’
Chimpanzee- U-shaped
A afarensis /africanus- U-shaped
Modern human- Parabolic
Their dental arcade is much larger than us.
Increase in size of molars are important.
Some important changes in the hands and wrists- more human like less apelike.
Facultative bipeds-
Obligate bipeds- the femus neck has a bigger dent thing and angle- makes a big difference in how legs work.
Afraensis and africanus have pelvis more like us, chimps have a bigger pelvis with huge range of motion compared to us. Ours have legs that swing forward and back, chimps they can go side to side.
Walking gait, chimps have awkward gait when bipedal, but ours is easy cuz of the forward back motion.
Laetoli footprints- differences in weight distro, but they had an almost modern foot and were likely obligate bipeds. It started with them then continued to later hominims.
Australopithecus africanus fossils
Taung Child a africanus.
“Mrs. Ples” a africanus- see traits you done see in other species. Appearance of bone structures related to chewing. Prominent canine pillars. Good candidate for a step on the branch leading to next species. Likely afrenses iwas ancestor of africanus.
A afarensis. fossils
“Lucy” – 3.2 mya – partial skeleton – Hadar, Ethiopia, A afarensis.
“Kadanuumuu” – 3.6 mya – adult male – north of Hadar region a afarensis.
“Selam” – 3.3 mya – mostly upper body – 3-year-old child – Hadar region A afarensis. Well preserved and helped shed new light on this species.
Afarensis upper body: evidence of tree climbing abilities- finger bones are a bit curved, means they were better at it than us. Scapula- similar to huma but greater range of arm movement, more like ape. Inner ear bones- keep balance. Apes have them shaped different for trees- Salems indicated adaptation to tree climbing.
Conclusions about A. afarensis and A. africanus
Obligate biped, but still spent a lot of time in the trees. Not as arboreal as others. Likely slept in trees and hid from predators. Could be a direct ancestor for us.
Paranthropus aethiopicus
2.5 mya Sagittal Crest Flaring cheek bone
the “Black Skull” discovered inkenya in 1985
410 cc brain.
Large molars
Masticatory apparatus.
Temporalis muscle, massaer muscle.
Saggital crest- on top of head.
Paranthropus boisei
2.3 to 1.2 mya East African Species
Very strong bite, more like gorilla.
Megadontia’
Possible version of tree:
Paranthropus robustus
2.0 to 1.0 mya South africa
120 cm and 35 kg for female
140cm and 50 kg for males.
Cranium differs them from the other species.
Smaller teeth than africanus.
Increase in molar size.
Strong mandible, powerful jaws.
520 cc brain. Big increase considering that they are still small bodied.
Drimolen 70 female no sagittal crest, 415 cc brain.
Drimolen 155
Male 450 cc brain, large sagittal crest
Sexual dimorphism.
Megadontia- mega sized teeth.
Paranthropines- seems to be a dead end- did not branch from there.. why?
Too specialized, the were not broadly adaptable. Die out when change happens
Miocene
end of Miocene 8-5 mya- this is when hominin appears and diverges from the other ape lineages.
- Origins of our genus Homo?
- Many researchers think Australopithecus afarensis is a direct ancestor to our genus homo- but no consensus and we cant be sure- currently no other reasonable options.
- Homo habilis
2.8–1.5 mya Mainly East Africa … maybe South Africa too? Currently appears to be the root of the genus tree. Koopbi Fora in Kenya.
* This means that homo hablis overlapped with Paranthropus and Australopithecus.
* The first fossils found are called ‘holotype’ fossils. Fossil # 7 in Oldavei George. (type fossil)
* Was controversial to assign this fossil to the genus homo.
* Cant tell male from female fossils- less sexual dimorphism than Australopithecus and paranthopenes.
* More similar to genus homo with small teeth and what not.
.
- Homo Rudophensis.
a proposed hominin species that is known from a few fossil specimens discovered in the Koobi Fora region near Lake Turkana in Kenya. The fossils date to around 1.9 million years ago. The taxonomic status of Homo rudolfensis is somewhat controversial, and it is considered by some researchers to be a variant or a different population of another hominin species, possibly Homo habilis.
Intermembral Index
Arm length/leg length x 100
- If your arms are half the length of your leg then the index is 50.
- Chimps = 105-110 arms are lightly longer than legs.
- Australopithecus/ Paranthropus= 85-95
- H. habilis= 80 – higher than us but lower than above.
- Homo sapiens= ≈70 interfemoral index.
- Presume as we became more efficient bipeds this evolved, less tree climbing
- Fossil 1813 from Koobi Fora, Kenya
cranial capacity of 510 cc. more than paranth. etc.
* contributes to our understanding of early human evolution and has been attributed to Homo habilis. This fossil offers insights into the brain size and morphology of early hominins,
- “Twiggy” Fossil 24 from Olduvai Gorge
1.8 mya- cranial capacity 590 cc. above average for the paranthopenes
* This fossil is believed to belong to Homo habilis and contributes to our understanding of the physical characteristics and brain sizes of early hominins. The relatively larger cranial capacity of this specimen indicates an advancement in brain size compared to earlier hominin species.
- Fossil 1470 from Koobi Fora
1.8 mya- missing all its teeth. 775 cc brain size.
* significant find attributed to Homo habilis. It’s characterized by having a relatively large cranial capacity compared to other fossils of similar age, marking a crucial point in the understanding of early human evolution. This fossil, dated to around 1.8 million years old, has a brain size of approximately 775 cc. Its brain size and other physical features align more closely with Homo habilis than other contemporary species, such as Paranthropus.
* Upper and lower jaws- 4th fossil. The lower jaw is not complete so able to reconstruct the other half since they are symmetrical. Both fossils are from Hadar but from different locations and different individuals.
- Hadar, Ethiopia
- very small teeth * more ‘modern’ looking than australopithecines and paranthopenes.
- The upper jaw is 2.3 mya and is the oldest fossil we found so far.
- likely corresponding to an ancient, transitional form possibly leading to the emergence of the Homo genus. The description of “more modern” features and smaller teeth could indicate a transitional stage between earlier australopithecines and the genus Homo. These fossils could represent a common ancestor or an early offshoot of the Homo genus, but they are not specifically identified as Homo habilis.
- 3 traits that distinguish Homo habilis from the Australopithecines and Paranthropines
made it clear that homo hablis was transitional to later homo species: homo erectus and homo egaster.
1) Increased Cranial Capacity- mean of 650 cc- more than 100 cc larger than paranthopenes who were larger than australopithecines. Did not increase in body size. Homo hablis degree of encephalization is greater than the aust and parant. Making it closer to where we ended up.
* Smaller Teeth Overall- more parabolic dental arcade. hablis has smaller molars and a parabolic dental arcade: U-shaped configuration of the dental arch in the mouth. It’s a more rounded dental arrangement compared to the rectangular or “V” shape observed in some other species. This dental arcade is characterized by a curve that is wider at the back and narrows towards the front of the mouth. This structure is notably found in some early humans like Homo habilis, distinguishing them from other hominin species with a different dental arrangement.Flattening of the face and reduced pragmatism.
2) More Advanced Precision Grip. Homo hablis has modern-looking hands. Power grip: used to hold a hammer- precision grip, more intricate for tool making. Distal phalanges – small fingertip bones. and apical tufts are more like us. Chimps have pointy apical tufts, ours are flat and wide. H hablis similar to ours. Paranthpos has slightly expanded aplicat tufts but not the austrolipithicans. Increased precision and dexterity- manufacture of stone tools- perhaps we will evolve back to having pointed tufts to aid in tapping tiny keys on our iPhones.
3) Very Modern-Looking Feet. The feet of aust and paranth are already similar to ours, homo hablis is more similar to us. Olduvai George 8- full h hablis- able to compare to modern foot. Compared to chimps and to us, we see an enlarged big toe designed for bipedal walking and carrying more weight.
- Trends in tooth size:
Ardipithecus ramidus and A. anamnesis had Smaller molars (like chimps and like us. )
* With Australopithecus aferensis we see a big jaw & big molars A. africanus: a big jaw & huge molars Paranthropus aethiopicus: huge jaw & megadontia
* Dead end p boisei and p robustus we see massive jaw & megadontia! Homo hablis around 3 million years ago smaller jaw & molars than afarensis (not far off chimps and us)
- Ardipithecus ramidus and A. anamnesis: These species had smaller molars, similar to both chimpanzees and early humans. They had a mix of characteristics that resemble both our common ancestors and some modern-day primates.
- Australopithecus afarensis: Displayed a larger jaw and bigger molars. This species was a likely ancestor of the Homo genus, including Homo habilis.
- Australopithecus africanus: Similar to A. afarensis with a big jaw and huge molars.
- Paranthropus aethiopicus: Exhibited an even more pronounced increase in jaw size and massive molars, termed “megadontia.”
- Paranthropus boisei and P. robustus: Both these species demonstrated a further increase in jaw size and megadontia. They are considered a separate branch from the Homo lineage and were evolutionary dead ends.
- Homo habilis: Around 3 million years ago, Homo habilis emerged with a smaller jaw and molars compared to afarensis, falling somewhere between the dental characteristics of chimpanzees and early humans. This is suggestive of a shift in diet and adaptations.
- The decrease in jaw and molar size among Homo habilis and the transition towards a more human-like dental pattern might signify dietary changes or adaptations that took place during that period. The shift from the larger, robust jaws and megadontia seen in Australopithecus and Paranthropus to the smaller, more modern dental characteristics in Homo habilis may coincide with changes in diet, tool use, or other adaptive behaviors
- Apical tufts
in the context of hand morphology, refer to the tip of the finger bones or phalanges. They are small projections found at the ends of the fingers. These tufts play a role in the mechanical support and functionality of the fingertip.
* In the case of Homo habilis, their apical tufts are considered to be more similar to those of modern humans rather than those of other earlier hominins. Chimps, for instance, have different-shaped apical tufts compared to humans. The differences in the apical tufts of various hominin species often reflect variations in their dexterity, grip, and tool-manipulating capabilities.
feet of Homo habilis
more similar to modern human feet compared to those of Australopithecus and Paranthropus. Specifically, Olduvai George 8, a full Homo habilis foot specimen, enabled comparisons to modern human feet. One significant feature is an enlarged big toe that facilitated bipedal walking and bearing more weight, which is a distinctive characteristic of human feet. This structural adaptation likely supported their mobility and stability while walking on two legs. * Fully adducted big toe:
- Fully adducted big toe:
means it’s parallel to other toes, unlike chimps.
abbducted big toe.
Diverges from the foot at an angle
- Human and H. habilis foot
Double arch- large heel bone. Longitudinal arch and transverse arch. Complex arch system means that their foot is mechanically set up for efficient weight transfers from the toes to the heel etc. important for obligate bipedalism