Chapter 9- Hearing, taste, and smell Flashcards
What is sound?
When we detect sound, we are actually detecting changes in air pressure that travel through the air to the ear (represented as a wave)
Amplitude
How big the wave is, measured as sound pressure. Amplitude is perceived as loudness, so a large amplitude= louder sound
Frequency
Measured in Hertz (Hz). How many cycles of up and down you have in one second. The more cycles you have, the higher the frequency and the higher the pitch (how frequency is perceived).
Which frequency can most people perceive?
Most people detect 20-20,000 Hz
What makes up the outer ear?
The pinna and ear canal
Pinna
The external part of the ear that you can see. Its function is to funnel sound waves into the ear canal. It also enhances certain frequencies- ridges on the outer ear amplify vibrations- the pitches of human speech especially. Sound waves propagate down the ear canal to the tympanic membrane.
Which structure marks the start of the middle ear?
The tympanic membrane
Which structures make up the middle ear?
The tympanic membrane and the ossicles
What is the function of the tympanic membrane?
Vibrations in the tympanic membrane at the same frequency as the sound, cause 3 ossicles (malleus, incus, and stapes) to move (also at the same frequency). Amplifies sound- vibration makes the sound louder
Which structures control the contact between ossicles?
Contact between ossicles is controlled by two muscles, the tensor tympani and stapedius muscles. This creates a protection mechanism from loud sounds that could be damaging- causes the ossicles to vibrate less. They also mute self made sounds (from inside the body)
3 ossicles
Stapes, incus, and malleus
Stapes function
Stapes (third ossicle) contacts oval window of the cochlea (with the same frequency). It transfers vibrations to 3 fluid filled canals in cochlea.
Cochlea structure
The cochlea makes up the inner ear. It is a spiral of 3 parallel fluid filled canals- the scala vestibuli, scala media, and scala tympani. The round window is a membrane separating the scala tympani from the middle ear. The basilar membrane and tectorial membrane separate the canals (scala vestibuli, media, and tympani).
Function of the basilar membrane
Sounds travel along the basilar membrane, but only parts of the basilar membrane will vibrate depending on the frequency of the sound. High frequencies vibrate the narrow, stiff base of the cochlea, and low frequencies vibrate the wide, floppy apex.
Organ of corti
A structure located on the basilar membrane of the cochlea. It contains inner and outer hair cells, which help to transduce sound waves into neural activity
Inner hair cells
IHCs detect the frequency of the sound
Outer hair cells
OHCs help discriminate between similar frequencies and assist with IHC signaling
Hair cells
Hair cells transduce sound waves into neuronal activity. They have 50-200 stereocilia (hairs) that will vibrate during sound transduction. Hair cells are not neurons and will not generate an action potential themselves- they synapse with auditory nerve fibers, which will generate an action potential to send to the brain
How do inner hair cells transduce sound?
- Vibrations of the basilar membrane bend stereocilia
- Tip links stretch and will open ion channels
- Calcium, potassium depolarize IHC. The depolarization does not trigger an action potential. If big enough, the depolarization will open voltage gated calcium channels and release NT
- Release NT onto auditory nerve (mainly Glutamate)
What can damage hair cells?
Loud, continuous noise like loud music damages the hair cells
Tonotopic organization
All levels of the auditory pathway are spatially arranged in a map according to the auditory frequencies they respond to. The organization begins in the cochlea, since frequency is ordered along the length of the basilar membrane. This is observed when the responses of auditory brain regions to tones of different frequencies are mapped. This is critical for sound localization.
Tuning curve
Auditory nerves have distinctive receptive fields or tuning curves. The lower the decibels, the more sensitive the auditory nerve is to the sound. Each auditory neuron responds to a specific frequency, but the neuron responds to a broader range of frequencies for more intense stimuli. A neuron is said to be “tuned” for the stimulus that evokes the greatest response.
Hair cells synapse onto
The vestibulocochlear nerve (the 8th cranial nerve)
What is the pathway of auditory information to the brain? (4)
- Hair cells synapse onto vestibulocochlear nerve (8th)
- 8th nerve synapses onto cochlear nucleus in brain stem. This is different from touch and pain- most information crosses to the other side, but some stays on the same side
- Info travels to both superior olivary nuclei
- Inferior colliculus (primary auditory centers of midbrain), then medial geniculate nucleus (thalamus), then auditory cortex