Chapter 7 – The Other Sensory Systems Flashcards
The intensity of a sound wave
Amplitude
Perception of the intensity of a sound
Loudness
A rapidly talking person sounds louder than slow music of the same physical amplitude
The number of cycles per second, measured in Hz
Frequency
The related aspect of perception of frequency
Pitch
Higher frequency sounds are higher in pitch.
With soundwaves, the height of each wave corresponds to ______, and the number of waves per second corresponds to _______
Amplitude; frequency
When it comes to structures of the ear, anatomists distinguish three parts:
The outer ear, the middle ear, and the inner ear
The outer ear structure of flesh and cartilage that sticks out from each side of the head
Pinna
By altering the reflections of sound waves, the pinna helps us locate the source of a sound. We have to learn to use that information because each person’s pinna is shaped differently from anyone else’s
The eardrum
Tympanic membrane
A membrane of the inner ear. The tympanic membrane connects to three tiny bones that transmit the vibrations to this area.
Oval window
The three tiny bones that transmit the vibrations of sound are known as:
Hammer, anvil, and stirrup
Latin names: malleus, incus, and stapes
Structure in the inner ear containing auditory receptors
Cochlea
A snail-shaped structure.
The auditory receptors that lie along the basilar membrane in the cochlea
Hair cells
Vibrations in the fluid of the cochlea displace the hair cells, thereby opening ion channels in its membrane. The hair cells excite the cells of the auditory nerve, which is part of the eighth cranial nerve
A cross-section through the cochlea, shows three long fluid-filled tunnels:
The scala vestibuli, scala media, and scala tympani
Concept that pitch perception depends on which part of the inner ear has cells with the greatest activity level
Place theory
According to this theory, each frequency activates the hair cells at only one place along the basilar membrane, and the nervous system distinguishes among frequencies based on which neurons respond.
The downfall of this theory is that the various parts of the basilar membrane are bound together too tightly for any part to resonate like a piano string.
Concept that the basilar membrane vibrates in synchrony with a sound, causing auditory nerve axons to produce action potentials at the same frequency
Frequency theory
For example, a sound at 50 Hz would cause 50 action potential’s per second in the auditory nerve.
The downfall of this theory in its simplest form is that the refractory period of a neuron falls far short of the highest frequencies we hear
Tenet that the auditory nerve as a whole produces volleys of impulses for sounds even though no individual axon approaches that frequency
Volley principle
For this principle to work, auditory cells must time their responses quite precisely, and the evidence says that they do. However, beyond about 4000 Hz, even staggered volleys of impulses can’t keep pace with the sound waves.
The hair cells along the basilar membrane have different properties based on their location, and they act as tuned resonators that vibrate only for sound waves of a particular frequency. The highest frequency sounds vibrate hair cells near the ______, and lower frequency sounds vibrate hair cells farther along the membrane near the ____
Base; apex
Impaired detection of frequency changes. For pitch perception, a fair number of people are not part of the normal distribution, includes an estimated 4% of people.
Amusia, often called “tone-deafness”
They have trouble recognizing tunes, can’t tell whether someone is singing off key, and do not detect a wrong note in a melody. Most people also have trouble singing even simple, familiar songs
Through which mechanism do we perceive low-frequency sounds, up to about 100 Hz?
At low frequencies, the basilar membrane vibrates in synchrony with the sound waves, and each responding axon in the auditory nerve sends one action potential per sound wave.
How do we perceive middle-frequency sounds, 100 to 4000 Hz?
At intermediate frequencies, no single axon fires an action potential for each sound wave, but different axons fire for different waves, and so a volley or group of axons fires for each wave
How do we perceive high-frequency sounds, above 4000 Hz?
At high frequencies, The sound causes maximum vibration for the hair cells at one location along the basilar membrane. High-frequency sounds excite hair cells near the base. Low-frequency sounds excite cells near the apex
What evidence suggests that absolute pitch depends on special experiences?
Absolute pitch occurs almost entirely among people who had early musical training and is also more common among people who speak tonal languages, which require greater attention to pitch
Area in the superior temporal cortex in which cells respond best to tones of a particular frequency
Primary auditory cortex or area A1
The organization of the auditory cortex strongly parallels that of the visual cortex.
The auditory cortex provides a kind of map of the sounds. Researchers call it a:
Tonotopic map
How is the auditory cortex like the visual cortex? 4
- Both vision and hearing have “what” and “where” pathways
- Areas in the superior temporal cortex analyze movement of both visual and auditory stimuli. Damage there can cause motion blindness or motion deafness.
- The visual cortex is essential for visual imagery, and the primary auditory cortex is essential for auditory imagery.
- Both the visual and auditory cortices need normal experience early in life to develop normal sensitivities.
What is one way in which the auditory and visual cortices differ?
Damage to the primary visual cortex leaves someone blind, but damage to the primary auditory cortex merely impairs perception of complex sounds without making the person deaf.
People with damage to the primary auditory cortex hear simple sounds reasonably well, unless the damage extends into sub cortical brain areas. Their main deficit is in the ability to recognize combinations or sequences of sounds, like music or speech.
Hearing loss that occurs if the bones of the middle ear fail to transmit sound waves properly to the cochlea
Conductive deafness or middle-ear deafness
Because people with conductive deafness have a normal cochlea and auditory nerves, they hear their own voices which can be conducted through the bones of the skull directly to the cochlea, bypassing the middle ear.
Hearing loss that results from damage to the cochlea, the hair cells, or the auditory nerve
Nerve deafness or inter-ear deafness
Can occur in any degree and may be confined to one part of the cochlea, in which case someone hear certain frequencies and not others.
Frequent or constant ringing in the ears
Tinnitus
Nerve deafness often produces tinnitus. In some cases, tinnitus is due to a phenomenon like phantom limb. Damaged part of the cochlea is like an amputation: if the brain no longer gets its normal input, axons representing other parts of the body may invade a brain area previously responsive to sounds, especially high frequency sounds
Which type of hearing loss would be more common among members of rock bands and why?
Nerve deafness is common among rock band members because their frequent exposure to loud noises causes damage to the cells of the ear
Describe the causes of nerve deafness
Can develop through inheritance or from a variety of disorders:
Exposure of the mother to rubella, syphilis, or other diseases or toxins during pregnancy; inadequate oxygen to the brain during birth; deficient activity of the thyroid gland; certain diseases, including multiple sclerosis and meningitis; childhood reactions to certain drugs, including aspirin; exposure to loud noises
Describe the causes of conductive deafness
Can be caused by diseases, infections, or tumourous bone growth. It is sometimes temporary, and if it persists, it can be corrected either by surgery or by hearing aids that amplify the stimulus.
Describe three methods of sound localization:
- Requires comparing the responses of the two ears. One cue for location is the difference in intensity between the ears. For high-frequency sounds, with wavelengths shorter than the width of the head, the head creates a sound shadow, making the sound louder for the closer ear.
- Difference in time of arrival at the two ears: sound coming from directly in front of you reaches both ears at once. A sound coming directly from the side reaches the closer ear about 600 µs before the other. Sounds coming from intermediate locations reach the two ears at delays between zero and 600 µs.
Most useful for localizing sounds with a sudden onset. - The phase difference between the ears: every sound wave has phases with two consecutive peaks 360° apart. If a sound originates to the side of the head, the sound wave strikes the two ears out of phase. How much out of phase depends on the frequency of the sound, the size of the head, and the direction of the sound. Useful for localizing sounds with frequencies up to about 1500 Hz in humans.
Which method of sound localization is more effective for an animal with a small head? Which is more effective for an animal with a large head? Why?
An animal with a small head localizes sounds mainly by differences in loudness because the ears are not far enough apart for differences in onset time to be very large. An animal with a large head localized sound mainly by differences in onset time because it’s ears are far apart and well-suited to noting differences in phase or onset time
Structures located in the vestibular organ, oriented in three planes and lined with hair cells; sensitive to the directional tilt of the head
Semicircular canals
Acceleration of the head at any angle causes the jelly like substance in one of these canals to push against the hair cells. Action potential’s initiated by cells of the vestibular system travel through part of the eighth cranial nerve to the brain stem and cerebellum.
Like the hearing receptors, the vestibular receptors are modified touch receptors. Calcium carbonate particles called _______ lie next to the hair cells. When the head tilts in different directions, they push against different sets of hair cells and excite them
Otoliths
People with damage to the vestibular system have trouble reading street signs while walking. Why?
The vestibular system enables the brain to shift eye movements to compensate for changes in head position. Without feedback about head position, a person would not be able to correct the eye movements, and the experience would be like watching a jiggling book page
Sensory network that monitors the surface of the body and it’s movements
Somatosensory system
Is not one sense but many, including discriminative touch, deep pressure, cold, warmth, pain, itch, tickle, and the position and movement of joints
Receptor that responds to a sudden displacement of the skin or high frequency vibrations on the skin
Pacinian corpuscle
A chemical, found in hot peppers, that produces a painful burning sensation by releasing substance P; high dosages damage pain receptors
Capsaicin
These somatosensory receptors respond to pain, warmth, and cold
Free nerve ending – unmyelinated or thinly myelinated axons
These somatosensory receptors respond to movement of hairs
Hair-follicle receptors
These somatosensory receptors respond to sudden displacement of skin; low frequency vibration or flutter
Meissner’s corpuscles
These somatosensory receptors respond to sudden displacement of skin; high-frequency vibration
Pacinian corpuscles