auditory perception Flashcards

1
Q

cause of sound

A

vibration of an object
movement alternately squeezes air molecules together and pulls them apart
creates a longitudinal pressure wave in air

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2
Q

function of pressure against time

A

high points - portions where pressure is high
air molecules squished together

low point - low pressure
air molecules pulled apart

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3
Q

amplitude

A

distance between baseline and peak of wave

amplitude used to derive intensity
loudness

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4
Q

decibels

A

logarithmic scale of relative intensities
reduces wide range of amplitudes to smaller scale

calculated with reference to our hearing threshold
used to determine loudness

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5
Q

sound intensity level

A

way of representing amplitude relative to a reference perception of loudness

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6
Q

period

A

time taken to complete one wavelength

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7
Q

frequency

A

number of periods per second

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8
Q

pitch

A

attribute in terms of which sound can be ordered on a musical scale

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9
Q

timbre

A

refers to quality which can make two sounds with the same pitch and loudness seem dissimilar
related to complexity

pure and complex tones

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10
Q

pure and complex tones

A

pure = a single frequency

complex = made of more than one frequency
can be broken down into individual pure tones

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11
Q

how do sounds have a clear pitch?

A

partials must be integers multiples of the fundamental frequency
called harmonics
if a sound has inharmonic partials it will be unpatched

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12
Q

outer ear

A

visible part of the ear - auricle
not vital for perception but has an effect
shape of ear important to perception of sounds

ear canal
extends down to eardrum (tympanic membrane)
resonant frequency = 1-5kHz

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13
Q

middle ear

A

two membranes joined by bones:
eardrum - tympanic membrane

ossicles - 3 tiny bones
hammer, anvil and stirrup

oval window - membrane like eardrum

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14
Q

why are the bones needed?

A

vibrations must travel from air to fluid
creates an impedance mismatch

harder for vibrations to move through fluid than air
middle ear helps to deal with it
= impedance matching device

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15
Q

function of the 2 membranes

A

eardrum bigger than oval window
power of vibrations concentrated into oval window
lever action of hinged bones
action amplifies strength of vibrations

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16
Q

inner ear

A
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17
Q

cochlea structure

A

snail shaped
two chamber separated by the cochlear partition
filled with perilymphic fluid

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18
Q

cochlea function

A

work as a frequency analyser
breaks incoming complex sounds down into pure tone components

also works as a transducer
converts mechanical energy at these different frequencies into electrical activity to travel to the brain

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19
Q

cochlear partition

A

splits cochlea in 2

basilar membrane
runs with cochlea
on top of the membrane in organ of Corti

organ of Corti contains
tectorial membrane - hinges over top of basilar membrane
hair cells (inner and outer) - topped with steriocillia (smaller hairs)

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20
Q

how vibrations move around the cochlear

A

vibrations flow from oval window through first chamber through helicotrema (gap at end) down to round window (another membrane) an reflected back

vibrations move around through cochlear

base end - end nearest middle ear
apex - tip of curled up formation

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21
Q

transduction process

A

basilar membrane moves in response to vibration in the perilymph
membrane vibrates at the same frequencies as the incoming sound

these vibrations bend the stereocilia on the inner hair cells against tectorial membrane

allows positively charged ions to enter cell
triggers release of neurotransmitters and an electrical signal is sent up auditory nerve to the brain

22
Q

how is pitch detected?

A

place and temporal coding

23
Q

place coding

A

at the base, basilar membrane is anchored, narrow and stiff
at apex, it is free, wide and loose

means it has different resonant frequencies at different points along length

points of maximum BM displacement = frequencies of incoming sound
(displacement where BM is vibrating most)

stimulate specific sets of inner hair cells
activates specific auditory nerve fibres
the tonotopy (place coding) is represented all the way up to auditory cortex

24
Q

temporal coding

A

BM moves in response to vibration in the perilymph
BM vibrates at the same frequencies as incoming sound

stereo cilia stimulated by peaks in BM vibration
means firing occurs at the same period of the incoming waveform
- known as phase locking

stereocillia stimulated at peak as hairs are brushed against tectorial membrane
- at point of maximum displacement

means firings happen at peaks of BM motion
so firing correspond to period of incoming waveform
- allows brain to detect the pitch of sounds

25
place coding at different frequencies
less reliable at low frequencies areas of vibration on BM bigger at lower frequencies less specific and precise encoding of pitch
26
temporal coding at different frequencies
breaks down at high frequencies not enough time for cells to recharge leads to temporal smearing peaks happening too close for coding to work firings overlap and brain doest know which firings correspond to which peak of waveform
27
function of inner hair cells
detect motion of basilar membrane ion flow causes electrical signals to brain
28
function of outer hair cells
amplify and sharpen motion of basilar membrane = cochlear tuning means we have good ability to discriminate different pitches ion flow causes mechanical changes expand and contract motions amplify and sharpen basilar membrane action = more precise
29
timbre coding
pitch represented by 2 mechanisms: place coding = where on the BM firing comes from temporal = when the firing is occurring timbre represented by which combinations of fibres are active at the same time
30
intensity coding
relies on the fact that there are low and high threshold auditory nerve fibres low threshold fibres discriminate quiet and moderate sounds - discriminate between low and medium amplitude sounds high threshold fibres kick in to discriminate moderate and loud sounds - discriminate between moderate and high amplitude sounds loudness of sound = total neural activity loud sound means all fibres are responding, quiet sounds mean only low responding
31
auditory pathway
starts with cochlear sent through auditory nerve to cochlear nucleus - acts as a relay station - send neural activity to other nuclei in brainstem then travels through superior olive - analyses location, where sounds are coming from - relies on precise timing so happens early then to inferior colliculus and medial geniculate - analyse pitch - relies on fairly precise timing, not quite as early as location processing ends in primary auditory area - analyses higher order features, such as timbre - less reliant on precise timing
32
auditory pathway simple
cochlear nucleus superior olive inferior colliculus medial geniculate primary auditory cortex
33
azimuthal plane
whether sound is coming from the left or right
34
intramural timing difference
sound from one side reaches that side first eg left reaches left first allows brain to detect direction sound is coming from brain very sensitive to these small differences in time
35
interaural level difference
if sound coming from left, right is shielded by the head sound arrives at far ear later arrives there quieter than other side when sound directly from side, ILD depends on the frequency
36
effect of frequency on ITD
to use ITDs, need to be able to match specific peaks in sound waves across both ears at higher pitch = shorter wavelength harder for brain to determine whether left or right came first misinterprets where peaks are relative to each other ITDs ambiguous or misleading at higher frequencies
37
effect of frequency on ILD
sounds diffract and bends around object smaller than its wavelength sound blocked by larger objects low frequencies diffract around the head - some sound reaches far ear - got there later but at a similar frequency high frequencies do not diffract - creates a head shadow - limited sound at the far ear - creates a larger ILD ILDs only useful at higher frequencies
38
how is elevation of sound detected?
ITDs and ILDs don't help pinnae do amplify some frequencies and reduce others create spectral cues by changing incoming frequency spectrum change the timbre complex waveforms represented as different component frequencies represented on a spectrum when sounds come from different heights shape of ears filter sounds in different ways - creates different timbres depending on elevation of noise = unique resonance patterns according to height
39
sounds from the front vs back
sound from directly in front or behind = ITD of 0 - takes just as long for sound to get to ears from front as back - equally as loud pinnae create small level differences we can also rotate our heads
40
neural coincidence model
axons transmitting electrical signals representing sound from both ears - acts as delay lines axons connected to neurons - act as neural coincidence detectors means neurons only fire when stimulated at same time by information from left and right ear measuring using ITD tuning curves look at different neurons and measure firing rate across different ITDs curve supports this model for some animals - terrestrial more likely to use opponent process analysis
41
how does the neural coincidence model work to determine angle of sound?
sound waves from straight ahead come in from both ears travels along until both left and right ears reach and feed into same neuron at same time - then fires this specific neuron during allows the brain to know that the sound in coming from directly in front if sound comes from right, hits right ear first signal comes in from right before left left stimulated later than right sounds reach neuron together causing it to fire different specific neruon fired - allows brain to determine that it is from an angle
42
opponent process analysis
involves 2 sets of broadly tuned neurons neurons tuned to left half of auditory space - in right hemisphere neurons tuned to right half of auditory space - in left hemisphere system calculates the difference between the two sets to work out where a sound is coming from - if most firing from left half, assume sound from left and vice versa - smaller differences mean closer to centre - no difference, assume sound straight infront
43
how are speech sounds made?
vibrations from the larynx travel upwards through vocal tract spectrum of sound is shaped by the articulators including: - soft palate, hard palate, tongue, teeth and lips and the resonant spaces (resonators) including: - chest, throat, mouth and nasal cavities larynx (voice box) contains folds of material air forced from lungs through fold, making them vibrate - vibrations create sounds
44
speech perception relies on more than the acoustic signal also reels on both top down (cognitive) and bottom up (perceptual) factor
45
fricatives
sounds made by forcing air through narrow gap in articulators eg s, sh, z a lot of energy through lots of frequencies but particularly in high frequencies shown by dark bands
46
vowel sounds
articulators can be used to shape our vocal tracts creates vowel sounds shown by stripy bands at lower frequencies peaks in spectrum - frequency components in vowel sounds that are particularly strong
47
coarticulation
the same sound is actually different depending on the acoustic context (neighbouring sounds) however we perceive these as being the same known as perceptual constancy reflects to down contribution of articulatory knowledge know what is needed to make these sounds so can remove small perceptual difference
48
McGurk effect
perceptual illusion visually presented sounds affect perception of the acoustic signal use of visual information inform interpretation of articulatory knowledge
49
linguistic knowledge
our perception of speech sounds is affected by the meaning of the context
50
phoneme restoration
if a familiar word is distorted (eg a missing sound) we put it back in without realising influence of lexical knowledge
51
Miller and Isard
perception of speech sounds is affected by the meaning od the context normal sentences grammatical string of words played people these sentences and asked to repeat them back - all sentences should have been equally intelligible but intelligibility decreased as stimuli became less meaningful or grammatical correct shows influence of semantic and syntactic knowledge - regardless of quality of acoustic inputs - contextual knowledge of linguistic factors essential for perception
52
sine wave speech
formants replaced with pure tones tracking the intensity modulations of those formants over time sine wave speech can be learnt has no fricative sounds etc only formants but due to formants and coarticulations can be learnt (formants influence perception of surrounding speech) shows influence of phonological, lexical and syntactic and semantic knowledge