Abiotic Stess Flashcards
Phases of stress
Alarm phase
Recovery phase
Hardening phase (acclimation)
Exhaustion phase
Adaption
A genetically determined level of resistance acquired by a process of selection of many genes
HD START protein
Homeodomain START transcription factor
Deeper more lateral roots, reduced stomatal density, higher abscisic acid, resistance to oxidative stress
Stress resistance mechanisms
Avoidance
Tolerance
Acclimation
Drought resistance mechanisms
Desiccation postponement (tissue hydration maintained) Desiccation tolerance (function while dehydrated) Drought escape (complete lifecycle in wet season)
Water deficit
Water content of a tissue or cell that is below the highest water content exhibited in the most hydrated state
Lines of defense against drought
Decreased leaf area
Deeper root growth
Stomatal closure (hydropassive/hydroactive)
Water deficit inhibits photosynthesis
Dehydrates mesophyll cells and impairs metabolism
Comparable solutes
Neutrally changed at physiological pH
Don’t interfere with enzyme function
Proline, glycine betaine
Cooling strats
Paraheliotropism/diheliotropism
Reflexive wax
Leaf size
ABA stress responsive genes
Osmoprotectants
Oxidative stress responses (peroxidase)
Movement of water and ions (aquaporins)
Membrane and protein stabilization (LEAs)
LEA proteins
Late embryogenesis proteins
Protects cell membrane (prevent crystallization of important molecules during desiccation)
Hydrophilic
Temperature compensation point
Temp at which amount of CO2 fixed by photosynthesis equals the amount of CO2 released by respiration in a given time
Signaling pathways for thermotolerance
ABA and SA signalling
Ethylene biosynthesis
Calcium signalling
Chilling injury
Disrupts starch conversion to sugars
Less CO2 exchange
Less photosynthesis
Destruction/degradation of chlorophyll
Chilling causes
Increased amount of desaturase enzymes
More unsaturated fats
Ice nucleation
Process by which water molecules form a stable ice crystal
Dehydration
Water moves from protoplast to extracellular ice
Antifreeze proteins
Bind to ice surface to slow or prevent more ice formation
Thermal hysteresis proteins
Sodicity
High Na+ content
Injures plants
Degrades soil structure
Salinity
Determined by electrical conductance
Osmotic effects of salt stress
Low osmotic potential
Less soil water potential
Salinity vs soil desiccation
Total amount of water available
Salt stress and ion toxicity primary effects
Na+, Cl-, SO4 2- accumulate in injurious concentrations
Deactivate enzymes
Plant membrane stability affected
Photosynthesis inhibited
Salt stress and ion toxicity secondary effects
Disruption of cell membrane integrity
Cell death
Salt reduction strats
Casparian strip - ion movement restriction into xylem Actively extrude Na+ into xylem Specialized salt glands on leaf surface Adjustment of water potential Changes in gene expression
Toxic trace element adaption
Exclusion
Internal tolerance
Hyperaccumulation (protection against herbivory)