Reproduction, hormones and plant hormones Flashcards

(47 cards)

1
Q

Asexual reproduction

A

one parent
No meiosis- only mitosis in an asexual life cycle
Offspring are genetically identical to each other and to the parent
No genetic variation is generated as existing gene combinations pass unchanged from generation to generation
Advantage: in an unchanging environment, well adapted parents produce well adapted offspring

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2
Q

Sexual reproduction

A

two parents (male and female)
Meiosis is part of every sexual life cycle
Offspring are genetically different from each other and from their parents
Genetic variation is generated as new gene combinations are produced in every generation
Advantage: species can evolve in a changing environment, as offspring can be better adapted than their parents

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3
Q

The role of meiosis in sexual reproduction

A

Meiosis and fusion of gametes have different roles

Fusion of gametes: a male and a female gamete join together to form a new individual. This brings alleles together in new combinations. Fusion of gametes is also referred to as fertilisation. It doubles the number of chromosomes each time it occurs.

Meiosis: one diploid nucleus divides to form four haploid nuclei. This breaks up parental combinations of genes, allowing new combinations to form when gametes fuse. By halving the number of chromosomes, meiosis reverses the doubling that is caused by the fusion of gametes, so there is no overall change in the number of chromosomes over a sexual life cycle

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4
Q

Male gametes

A

Motility of gamete: male gametes travel to the female gamete
Size of gamete: smaller, allowing faster movement
food reserves: less- only enough for the gamete
numbers produced: more- often very large numbers

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5
Q

Female gametes

A

Motility of gamete: female gametes are non-motile
Size of gamete: larger due to stores of food reserves
Food reserves: more- for embryo development
Numbers produced: few- sometimes only one

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6
Q

The male reproductive system

A
  • erectile tissue- fills with blood to enlarge and harden the penis
  • penis- penetrates the vagina so semen can be ejaculated near the cervix
  • urethra- transfers semen during ejaculation and urine during urination
  • testis- produces sperm and testosterone
  • sperm duct- transfers sperm during ejaculation
  • seminal vesicle/prostate gland- secrete alkaline fluid, proteins and fructose which is added to sperm to make semen
  • epididymis- stores sperm until ejaculation
  • scrotum- holds testes at lower than core body temperature to promote sperm development
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7
Q

The female reproductive system

A
  • ovary- produces eggs, oestradiol and progesterone
  • uterus- provides for the needs of the embryo and then the foetus during pregnancy (protection, supply of food and oxygen, and removal of waste products)
  • vulva- protects internal parts of the female reproductive system
  • oviduct- collects eggs during ovulation, and provides a site for fertilisation and then moves the embryo to the uterus
  • cervix- protects the foetus during pregnancy and then dilates to provide a birth canal
  • vagina- stimulates the penis to cause ejaculation and forms the birth canal
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8
Q

The menstrual cycle

A

The menstrual cycle takes about 28 days and is composed of cycles that happen in the ovaries and uterus.

In the ovarian cycle, follicles start to develop in the ovary, but usually only one completes development and releases its egg on about day 14. After ovulation, the wall of the follicle enlarges and becomes the corpus luteum. If an embryo isn’t present, the corpus luteum breaks down towards the end of the menstrual cycle.

In the uterine cycle, the endometrium becomes thickened, with enhanced blood supply, in preparation for implantation of an embryo. If no embryo is present, the thickening breaks down and passes out of the body. This is menstruation (”period”). As the start of the menstruation is an obvious event, it is used to define day one of the cycle

Four hormones regulate the menstrual cycle by negative feedback and positive feedback. FSH and LH bind to receptors in the membranes of follicle cells. Oestradiol and progesterone affect gene expression and therefore development in the uterus and other parts of the female body.

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9
Q

The role of protein hormones produced by the pituitary gland (menstrual cycle)

A
  • FSH- rises to high levels in the first ten days of the menstrual cycle. During this time it stimulates the development of follicles, each containing an oocyte (egg) and follicular fluid. FSH also stimulates oestradiol secretion by the wall of the developing follicle
  • LH rises to a sudden sharp peak on about day 14. It stimulates the maturation of the oocyte and ovulation by bursting of the follicle wall. LH then promotes the development of the corpus luteum, which secretes oestradiol (positive feedback) and progesterone
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10
Q

Role of steroid hormones produced by the follicle wall and corpus luteum in the menstrual cycle

A
  • oestradiol rises to a peak in the second week of the cycle. It stimulates the repair and thickening of the endometrium after menstruation and an increase in FSH receptors to make the follicles more receptive to FSH, which boosts oestradiol production (positive feedback). When it reaches high levels, oestradiol inhibits FSH secretion (negative feedback) and stimulates LH secretion
  • Progesterone levels rise following ovulation, reaching a peak and then dropping back to a low level by the end of the menstrual cycle if no embryo is present. Progesterone promotes the thickening and maintenance of the endometrium. It also inhibits FSH and LH secretion by the pituitary gland (negative feedback)
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11
Q

The process of fertilisation (fusion of sperm and egg cell)

A

Fertilisation is the fusion of a sperm with an egg to form a zygote. Sperm have receptors in their plasma membrane to detect chemicals released by the egg. This enables the sperm to swim towards the egg. Around the egg is a cloud of follicle cells and a layer of glycoproteins (zona pellucida)

The sperm push between the cells and digest a route through the glycoproteins to reach the plasma membrane of the egg cell. The sperms plasma membrane has proteins that bind to the egg cell’s plasma membrane. The first sperm that manages to reach the egg binds to it and the membranes of the sperm and egg fuse together. The sperm nucleus then enters the egg cell. This is the moment of fertilisation. Immediately afterwards the layer of glycoprotein hardens to prevent further sperm entry

The sperm tail does not penetrate the egg during fertilisation or is broken down inside the zygote. Sperm mitochondria may also penetrate, but they are destroyed by the egg cell.

The nuclei from the sperm and egg remain separate until the zygotes first mitosis. Then both nuclear membranes break down, releasing 23 chromosomes from each nucleus. These chromosomes, half from the mother and half from the father, participate jointly in mitosis, using the same spindle of microtubules. Two genetically identical nuclei are produced, each with 46 chromosomes.

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12
Q

Why do male and female gametes need to be differently adapted

A

Male gametes travel to female gametes and because of this they have very different adaptations. Male gametes in all animals and some plants are motile- they can swim. The faster they swim, the more chance of reaching the egg and fertilising it, so small size and an efficient propulsion system are needed. Male gametes are produced in larger numbers than female, to increase the chance of one of them fertilising an egg.

In humans (and other species) female gametes are much larger as they contain nearly all the food reserves for the early development of the embryo. Because of the large investment of resources, smaller numbers of female than male gametes are produced. Female gametes have a mechanism for allowing one sperm to penetrate, but not more.

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13
Q

Ovum adaptations

A
  • haploid nucleus- contains the 23 chromosomes that are passed from mother to offspring
  • two centrioles
  • first polar body- not needed so breaks down
  • plasma membrane
  • layer of follicle cells (corona radiata)
  • zona pellucida- protects the egg cell and restricts the entry of sperm
  • cortical granules- harden the zona pellucida to prevent multiple fertilisation (polyspermy)
  • cytoplasm/ yolk- contains droplets of fat and other nutrients needed during early stages of embryonic development
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14
Q

Sperm (spermatozoa) adaptations

A
  • acrosome- contains enzymes which digest the zona pellucida around the egg
  • plasma membrane
  • centriole
  • mid-peice with helical mitochondria- produce ATP by aerobic respiration to supply energy for swimming and other processes in the sperm
  • haploid nucleus- contains the 23 chromosomes that are passed from father to offspring
  • tail/flagella- provides the propulsion that allows the sperm to swim up the vagina, uterus and oviduct until it reaches the egg
  • protein fibres to strengthen the tail
  • microtubules in a 9+2 array- make sperm tail beat from side to side and generate the forces which propel the sperm
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15
Q

Barriers to prevent polyspermy (Multiple sperm fusing with the egg)

A

Fusion of more than one sperm with an egg is called polyspermy. Structure in the sperm and the egg make polyspermy very infrequent. A coat of glycoproteins (zona pellucida) surrounds and protects the egg. When sperm make contact with the zona pellucida, they release the contents of their acrosome. This is the acrosome reaction. Enzymes from the acrosome digest a small region of the zona pellucida, allowing the sperm to penetrate and reach the egg.

When the first sperm has fused with the egg, the many cortical granules (vesicles) near the eggs plasma membrane release their contents by exocytosis. This is the cortical reaction. The enzymes that are released toughen the zona pellucida making it very difficult for any more sperm to penetrate it. The enzymes also change specific glycoproteins in the zona pellucida to which sperm bind, so that this can no longer happen.

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16
Q

what is IVF?

A

Natural fertilisation is in vivo, as it occurs inside the body. Fertilisation can also happen outside the body in dishes in the laboratory. This is in vitro fertilisation, usually abbreviated to IVF. The procedure has been used extensively to overcome male/ female fertility problems. There are various protocols for IVF.

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17
Q

What are the stages of IVF

A
  1. down-regulation: every day for two weeks, the woman has an injection or nasal spray containing a drug to stop the pituitary gland from secreting LH or FSH. Secretion of oestradiol and progesterone therefore also stops, pausing the womans normal menstrual cycle, allowing doctors to control the timing and amount of egg production
  2. FSH injections- intramuscular injections of FSH are given daily for 7-12 days, to stimulate follicles to develop. The aim is to generate very high FSH concentration, resulting in far more follicles than normal (superovulation). The aim is 8-15 follicles, each containing an egg.
  3. hCG injection- when the follicles are 18mm in diameter, they are stimulated to mature by an injection of human chorionic gonadotropin (hCG). This is the hormone embryos secrete to signal to their mother that they are present
  4. Egg collection- this is a minor surgical procedure that takes around 20 minutes. A micropipette mounted on an ultrasound scanner is used to draw the eggs out of the follicles. Egg collection is done about a day and a half after hCG injection.
  5. Fertilisation- each egg is mixed with 50,000-100,000 sperm cells in sterile conditions in a shallow dish, which is incubated at 37 degrees celsius until the next day
  6. Embryo transfer- one or more embryos are placed in the uterus when they are about 48 hours old. A progesterone tablet is placed in the vagina, to ensure that the uterus lining is maintained. If the embryos implant and continue to grow, the pregnancy that follows is no different to a normal pregnancy that began by natural conception.
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18
Q

What is hCG (human chorionic gonadotropin)?

A

The hormone human chorionic gonadotropin (hCG) is a medium- sized protein. hCG is produced by the trophoblast cells in the blastocyst and subsequent embryonic stages, so it is an early indicator of the presence of an embryo. Later in pregnancy, cells in the planceta continue the secretion of hCG.

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19
Q

How do pregnancy tests work?

A

Tests for pregnancy are based on the detection of hCG in a womans urine. A test stick has an absorbent strip inside a plastic casing. There are three types of monoclonal antibody in the strip.

A urine sample is placed on the strip where it sticks out at one end. The urine is drawn through the strip by capillary action. it first meets antibody A and has a molecule of blue dye detached. Any hCG molecules in the urine bind to this antibody. Antibody A with or without bound hCG, moves on with the urine along the strip. Next is a band of immobilised antibody B which binds to antibody A which have hCG bound, creating a blue band which indicates pregnancy. Molecules of antibody A without hCG move along the strip and reach a band of IgG antibody, to which they bind, creating a blue band which indicates the test has worked.

20
Q

What is gametogenesis?

A

Gameteogenesis is the production of gametes. Male gametes are sperm (spermatozoa) and female gametes are eggs (oocytes). Spermatogenesis happens in the testes and oogenesis in the ovaries. The four stages in both types of gametogenesis are mitosis, cell growth, two divisions of meiosis and then differentiation.

21
Q

What is spermatogenesis?

A

Spermatogenesis occurs in the seminiferous tubules of the testes, which are lined with a germinal epithelium.
Spermatogonia (diploid germ cells) divide by mitosis to produce more spermatogonia.
Some spermatogonia grow into primary spermatocytes, which undergo meiosis I to form two secondary spermatocytes.
Secondary spermatocytes undergo meiosis II to form four haploid spermatids.
Sertoli cells provide nourishment and support during development.
Spermatids undergo differentiation (spermiogenesis) to become mature spermatozoa.
The developing sperm cells are gradually pushed toward the lumen of the seminiferous tubule.
Leydig (interstitial) cells, located between tubules, produce testosterone, which stimulates spermatogenesis.

22
Q

What is oogenesis?

A

Oogenesis occurs in the ovaries, starting in the germinal epithelium. During foetal development, these cells divide by mitosis and migrate into the cortex. By 4–5 months, they grow and start meiosis, but pause in prophase I. At birth, a female has primary follicles, each containing a cell arrested in meiosis I.

At puberty, during each menstrual cycle, some primary follicles begin development, but usually only one matures and ovulates. Meiosis I completes, producing a large secondary oocyte and a small first polar body (which degenerates). The secondary oocyte begins meiosis II but pauses in metaphase II. Meiosis II only completes if fertilization occurs, forming an ovum and a second polar body (which also degenerates).

23
Q

Development of a blastocyst

A

The zygote produced by fertilisation in the oviduct divides by mitosis to form a 2-cell embryo. Further rounds of the cell cycle double the number of cells about every 18 hours. Because the egg has a large amount of cytoplasm, these early rounds of division can happen without any cell growth, so cell size decreases. Initially the embryo is a solid ball of cells. When it is 6 or 7 days old, it has changed into a hollow ball, due to unequal divisions and cell migration. This embryonic stage is the blastocyst. It has about 250 cells and is 200 micrometres in diameter.

While the embryo is developing it is wafted down to the oviduct by cilia in the oviduct wall. The embryo arrives in the uterus when it is about 7 days old and has become a blastocyst. The embryo has used up the food reserves of the egg by this stage, so it must obtain supplies from its mother. Cells in the outer layer (trophoblast) of the blastocyst secrete enzymes that loosen the connections between cells in the lining of the uterus wall (endometrium). This allows the blastocyst to penetrate the uterus wall and become implanted. If implantation does not occur, the embryo is not supplied with enough food and the pregnancy fails.

24
Q

What is the placenta?

A

The placenta is a disc-shaped structure, embedded in the uterus wall. It’s development begins about four weeks after conception in humans. The basic functional unit is a finger-like piece of foetal tissue called a placental villus. The size of the placenta and the number of villi grow larger during the pregnancy, so that the surface area matches the increasing demands of the foetus for exchange of materials with the mother. Maternal blood flows in spaces around the villi (intervillous spaces). This is a very unusual type of circulation as elsewhere blood is almost always retained in blood vessels. Foetal blood is pumped to the placenta through arteries in the umbilical corod by the foetal heart and then passes through a dense network of blood capillaries close to the surface of each villus.

25
The role of the placenta in transfer of substances?
The distance between foetal and maternal blood is very small, as little as 5 micrometres. The cells that separate the maternal and foetal blood form the placental barrier. This must be selectively permeable, allowing some substances to pass but not others. Glucose, amino acids, oxygen and all other substances required by the foetus pass from maternal to foetal blood. Carbon dioxide, urea and other waste products pass from foetal to maternal blood. Diffusion, facilitated diffusion, osmosis and endocytosis are all used in these exchanges for specific substances. The blood passes from the placenta to the body of the foetus in the umbilical vein. The foetus can be retained in the uterus to a much later stage of development in placental mammals than in mammals such as kangaroos which don’t develop a placenta.
26
What is puberty?
Puberty is development from childhood to sexual maturity. When the brain determines that the time for puberty has arrived, the peptide hormones GnRH (gonadotropin-releasing hormone), is synthesised by neurons into the bloodstream. Secretion continues through puberty and adulthood. GnRH stimulates secretion of FSH and LH by the pituitary gland. These two hormones are gonadotropins because they cause changes in the gonads- testes and ovaries.
27
Role of LH and FSH in boys
In boys, FSH stimulates testis growth and LH stimulates secretion of the steroid hormone testosterone from the testes. Testosterone causes the development of the male secondary sexual characteristics in boys during puberty such as enlargement of the penis, growth of pubic hair and deepening of the voice due to growth of the larynx.
28
Role of LH and FSH in girls
In girls, FSH stimulates the development of follicles in the ovary. The follicle wall secretes the steroid hormone oestradiol. LH causes the development of the follicle wall into the corpus luteum after ovulation, which continues the secretion of oestradiol. Oestradiol causes the development of secondary sexual characteristics in females during puberty such as enlargement of the uterus, development of the breasts and growth of pubic and underarm hair.
29
Hormones of pregnancy and childbirth
Secretion of hCG throughout pregnancy stimulates continuing secretion of oestradiol and progesterone. These two steroid hormones are initially secreted by the corpus luteum, but after about 10 weeks of pregnancy the placenta takes over. They are essential to prevent miscarriage and allow the pregnancy to continue. Oestradiol and progesterone prevent degeneration of the uterus lining. Progesterone prevents contractions of the muscular outer wall of the uterus (myometrium) by inhibiting oxytocin secretion by the pituitary gland. Concentrations of both steroid hormones gradually rise, but near the end of a pregnancy the foetus signals to the placenta to stop secreting progesterone, so the concentration starts to decline. This allows the pituitary gland to start secreting oxytocin. Oxytocin stimulates contraction of the muscles in the myometrium. The contractions are detected by stretch receptors, which signal to the pituitary gland to increase oxytocin secretion (positive feedback). Increased oxytocin concentration makes the contractions more frequent and more vigorous, causing more oxytocin secretion. This is an example of positive feedback feedback. In this case, it has the advantage of causing a gradual increase in the myometrial contractions, allowing the baby to be born with the minimum intensity of contraction. The baby is pushed out through the cervix and vagina. The umbilical cord is broken and the baby takes its first breath and achieves physiological independence from its mother.
30
Role of hormone replacement therapy in menopause
Hormone replacement therapy (HRT) is used to relieve menopausal symptoms. It supplements levels of the oestradiol and progesterone that naturally decrease as a woman approaches the menopause. HRT can relieve symptoms such as hot flushes, night sweats, mood swings, vaginal dryness and reduced sex drive. Many of these symptoms pass after a few years, but they can be unpleasant and HRT offers relief to many women. It can also reduce osteoporosis (loss of bone density) which is more common after the menopause.
31
Male and female parts of a flower
- Anthers produce pollen containing male gametes. Diploid cells inside the anther divide by meiosis to produce four haploid cells, each of which develops into a pollen grain. The nucleus inside a pollen grain divides by mitosis to produce the male gametes. Pollen grains develop a thickened wall for protection during pollination - Ovules are ovoid structures inside the ovary. One cell in the centre of the ovule grows particularly large and then divides by meiosis. One of the haploid nuclei produced divides three times by mitosis to produce eight haploid nuclei, one of which becomes the female gamete (egg nucleus).
32
pollination, fertilisation and embryo development
- Pollination is the transfer of pollen from an another to a stigma. The pollen is usually moved by wind or by animals - Fertilisation happens inside an ovule. Each pollen grain that lands on the stigma grows a tube d own the style to the ovary, to transport the male gametes from the pollen grain. Once inside the ovary, the pollen tube grows to one ovule and digests a route into it. When the tube reaches the centre of the ovule where the female gamete (egg) is located, the male gametes are released and fertilisation occurs, resulting in the production of a zygote. - Embryo development- the zygote divides repeatedly by mitosis and the cells produced develop into an embryo, with an embryo root, embryo shoot and either one or two embryo leaves (cotyledons). The outer layers of the ovule develop into the seed coat.
33
What type of reproduction occurs in flowers?
The type of reproduction that happens inside flowers is sexual. Many flowers are hermaphroditic because they have both male and female parts, so can act as both a male and female parent. In some cases, hermaphrodite plants can act as both the male and female parent at the same time, by self-pollination and self-fertilisation. Even so, this is not asexual reproduction as all the key events of sexual reproduction happen: meiosis, gamete production and fertilisation.
34
How can the chances of cross pollination be increased?
Cross-pollination is the transfer of pollen from the anther of one plant to the stigma of another. An outside agent is needed to transfer the pollen, usually an animal or the wind. Cross-pollination leads to fusion of male and female gametes from different plants, so promotes genetic variation and therefore evolution, which is essential in an ever changing environment. Many plants are hermaphroditic; they produce both pollen with male gametes and ovules with female gametes, making self pollination possible, causing inbreeding. In most human societies there are taboos against closely related individuals reproducing. This is because of high rates of miscarriage and genetic disease in offspring in such cases. Inbreeding causes the chance of a rare recessive allele in one ancestor being inherited twice by an individual, thus causing a genetic disorder. The offspring of two genetically identical plants tend to be healthy and grow strongly (hybrid vigour). Natural selection therefore tends to favour plants that reproduce by cross pollination, and mechanisms have evolved to promote it and reduce the change of inbreeding by self pollination. Some methods include: separate male and female plants so athers and carpels are on different plants (Ginkgo biloba or urtica dioica) separate male and female flowers on the same plant so anthers and carpels arent in the same flower (zea mays or betula papyifera) stigmas and pollen on anthers mature at different times (protandry = pollen first, protogyny = stigma first) (Nelumbo nucifera, digitalis purpuera)
35
Structure of insect pollinated flowers?
- Petals- large and brightly coloured to advertise the flower, act as a landing stage and guide the insects movements in relation to the position of the anthers and stigma. Petals in many insect-pollinated plants release a scent to allow insects to find the flower more easily - Pollen grains- spiky so they stick to the anther initially, and then to the visiting insect. The grains can be relatively large as they do not need to be light to be blown by wind. - Anther- positioned to deposit pollen on the insect - Stigma- positioned where a visiting insect will brush past it and sticky so pollen from it adheres. - Nectaries- glands that secrete nectar; nectaries are positioned deep inside the flower so insects can only reach them by brushing past the anthers and stigma
36
Self incompatibility mechanisms
Despite adaptations that reduce the chance of self-pollination the stigma of hermaphrodite plants may receive pollen from that plants own anthers. In many plants this pollen fails to germinate, or the pollen tube. stops growing before it reaches the ovary. This protection against inbreeding due to a single plant acting as both male and female parent is self-incompatibility. It has evolved more than once and there are different mechanisms, but there is always a genetic bias, with alternative alleles of one or more genes. Plants with the same self incompatibility alleles cannot produce offspring together.
37
Seed dispersal and germination
Seeds are often transported long distances from the parent plant. This is seed dispersal. It helps to spread the species and reduces competition between offspring and parent. Seed dispersal and pollination are separate processes in the sexual life cycle of plants. Seed dispersal is the function of the fruit, which develops from the ovary of the flower. The structure of fruits is very variable and corresponds to the seed dispersal strategy: - fleshy and attractive for animals to eat - feathery or winged to catch the wind - covered in hooks that catch onto the coats of animals - explosive seed germination follows dispersal - it happens when the conditions are suitable. There must be water, oxygen and warmth - food reserves inside the seed are mobilised by being digested and transferred to the growing embryo; the embryo shoot grows upwards and the embryo root downwards.
38
Trophic responses in seedlings
Plants control the direction of growth of their roots and shoots. These parts of the plant have meristems at their tips, where cell division and enlargement occur. Roots and their shoots become curved if one side grows more quickly than the other side. This type of unequal growth happens in response to external stimuli perceived by the plant, such as the direction of gravity or sunlight. Differential growth responses to directional stimuli are called trophic responses or tropisms. - positive tropism is growth towards the stimulus - negative tropism is growth away from the stimulus Most roots are positively graviotropic (geotropic). This means they grow downwards, in the same direction as gravity. Most shoots are positively phototropic and negatively gravitropic. This means they grow towards the source of light and, in darkness, they grow upwards in the opposite direction of gravity. The roots and shoots of plants do not all show the same tropic responses. You can investigate these responses qualitatively- by recording observations in drawings or photos- or quantitatively. The angle of curvature of a root or shoot that has carried out a trophic response is an example of a quantitative measure.
39
What is a positive phototropism?
If one side of a shoot receives a higher intensity of light than the opposite side, the shoot responds with positive phototropism. The response to lateral light is differential growth, with the sunnier side growing more slowly than the shadier side. When the shoot has curved towards the direction of maximum light intensity, growth becomes equal on all sides, so the shoot carries on growth in that direction. The benefit of positive phototropism is obvious: it increases the amount of light absorbed by a shoots leaves for use in photosynthesis. This is particularly important where there is competition with other plants for light and where the brightest light is to one side rather than directly above. A pigment detects light intensities in the shoot tip and auxin (growth promoting phytohormone) is then redistributed in the shoot, causing differential growth.
40
what are phytohormones?
A hormone is a chemical message that is produced and released in one part of an organism to have an effect in another part of the organism. Plants and animals produce, transport and use hormones in different ways, so plant hormones are called phytohormones. A variety of chemicals rare used as phytohormones by plants, with roles in the control of growth, development and responses to stimuli.
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What are the roles of phytohormones?
1. Growth Phytohormones. can promote or inhibit growth, by affecting rates of cell division and cell enlargement. For example, gibberellin promotes stem growth. Wheat and other crop plants have been bred with short stems by introducing alleles which make the plant less responsive to gibberellin. 1. Development Phytohormones can promote or inhibit aspects of development- for example, whether a but starts to grow to produce a side shoot, or whether the apex of a stem produces more leaves or changes to produce flowers. Fruit ripening is promoted by the phytohormone ethylene in many plants. 1. Responses to stimuli Tropic responses are controlled by phytohormones. Tendrils of climbing plants respond to touch stimuli by coiling around a potential support. Communication using electrical signals is used for rapid responses, such as the capture of an insect by a Venus flytrap plant. However, a phytohormone called jasmonic acid triggers the subsequent secretion of enzymes to digest the fly.
42
What are auxin efflux carriers?
Auxin can enter cells by passive diffusion if its carboyxl group (COOH) is uncharged. The cytoplasm of plant cells is alkaline, so auxin is deprotonated (H+) from its carboxyl group, and becomes negatively charged (COO-). This traps the auxin inside the cell. However, auxin efflux carriers can pump negatively charged auxin molecules across the plasma membrane into the surrounding cell wall. The cell wall is acidic, so the auxin reverts to its uncharged state. It can then diffuse into an adjacent cell. To transport auxin across a tissue, efflux carriers are moved to the same end of each cell. Auxin is therefore pumped in the same direction by all the cells in the tissue, generating a concentration gradient.
43
what is the acid growth theory?
Auxin (a plant hormone) promotes the synthesis of proton pumps (H+-ATPases) within the cell. These proton pumps are inserted into the plasma membrane of plant cells. The pumps actively transport H+ ions from the cytoplasm into the cell wall, resulting in acidification of the cell wall. This acidic environment weakens the hydrogen bonds between the cellulose microfibrils in the cell wall, a process that loosens the structure of the wall. As a result, water enters the cell by osmosis due to the lower osmotic potential in the vacuole, causing the cell to swell and expand. The swelling of the cell wall allows for cell elongation and growth.
44
Interactions between auxin and cytokinin
- Auxin is produced in shoot tips and transported in the phloem down stems and into roots. - Cytokinin, another type of phytohormone, is produced in root tips and transported in xylem up roots and into stems. The amounts of auxin and cytokinin produced in a plant provide a means of balancing root and stem growth. They work together to stimulate some processes (synergism) and have opposite effects in other cases (antagonism) Auxin produced by the shoot tip inhibits growth of axillary buds nearby, so the main shoot grows strongly, without competition from side branches. If the main shoot of a plant is eaten by an animal, auxin concentrations decrease, which allows one or more nearby axillary buds to develop into branches of the stem, replacing the lost main shoot. These side shoots produce auxin, which stimulates the root growth needed to supply water and nutrients and inhibits development of more axillary buds.
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Syngergism between auxin and cytokinin
cell division in root and shoot tip meristems (synergism) cell growth in root and shoot tip meristems (synergism) both auxin and cytokinin promote these action
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Antagonism between auxin and cytokinin
development of new roots and branching of roots (antagonism): this is promoted by auxin and inhibited by cytokinin branching of stems by the growth of axillary buds (antagonism): this is inhibited by auxin and promoted by cytokinin
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Role of ethylene (ethene) in fruit ripening)
Succulent fruits such as peaches ripen when the seeds inside are fully developed. Animals are then attracted to eat the fruits and disperse the seeds. During fruit ripening, the flesh becomes softer, acids and starch are converted to sugar and the skin colour changes and volatile scents are produced to advertise the fruit to animals. Many fruits are stimulated to ripen by ethylene (ethene) and produce this phytohormone as they ripen. so there is a positive feedback mechanism that promotes rapid ripening. Ethylene is volatile, so can diffuse through the air from ripening fruits to other fruits, initiating their ripening. This helps to synchronise the ripening of fruits, increasing the attractiveness of a plant with fruits to animals and encouraging the dispersal of the seeds.