plants and water Flashcards
drought
- threatens agricultural livelihoods and causes food insecurity
- e.g. Afghanistan 2018, caused large displacement, exacerbated migrant crises etc
roots
- absorptive interface with soil
- lag on research on rooting behaviour, hormone signalling etc
- structure studied using CT scanning
root structure
- Dicots, large fleshy primary tap root, branches of first-order lateral roots, may branch again into second-order lateral roots
- Monocots, primary seminal root branches into first and second order lateral roots, tend to be more branched, crown roots just under soil surface
- root system architecture adapted to environment, low nutrients leads to more branched lateral roots
root hairs
- increases surface area for absorption
- elongations from single epidermal cell, vacuole pushes cell membrane and wall outwards
- interactions of hormones determines number (depends on environmental conditions), distribution etc
- relatively short lived, continuously regenerated
- specialised, thin permeable cell walls
symbiotic root fungi, ectomycorrhizal
- 80% plant species
- mantle of fungal cells coat outer root surface, roots do not directly interact with soil
- enhances water and nutrient exchange
- increases protection
- Hartig net, fungi can penetrate through cell walls (not into cells through membrane)
- fungi extracts some carbon from plant
- increases surface area through mycelia
root tissues
- cortex layer stores nutrients, water, carbohydrates
- vascular tissue bounded by endodermis
- phloem, mostly for transport of carbohydrate down to roots
- xylem
endodermal layer
- specialised lignified cells
- Caspian strip waterproofed by suberin encloses cell wall
- water in apoplastic route (in channels within cell walls) forced into symplastic route (into cells)
- allows cellular control of water entering vasculature (removal of bacteria, viruses etc)
plant mineral deficiencies
- plants need 16 or more nutrients (debated) taken up from soil
- leaf discolouration can diagnose deficiencies
- mobile nutrients include N, P, K, Mg
- able to move out of older leaves to younger plant parts when supplies are inadequate, older leaves are less efficient as they are at bottom of plant, apoptosis often triggered
- immobile nutrients include Mn, S, Cu, Fe, Ca, B
- plants struggle to move them elsewhere, deficiency symptoms first appear in new growth or young leaves and can be localised
human mineral deficiencies
- all plant derived and therefore soil derived, problem with intensive agriculture depleting nutrients, can rotate crops/ supplement soil
- undernutrition (not receiving enough calories or correct balance of nutrients) accounts for 45% worldwide child death
- vitamin D deficiency a problem in UK
- vitamin A deficiency a problem in other countries, rice lacks vitamin A, can lead to disability and death (blindness)
- GMO ‘golden rice’ is fortified with vit A
dicot stem xylem cells and tissues
- vascular bundles in circular arrangement around pith
- as the stem matures, secondary growth occurs through the vascular cambium
- produces additional xylem cells (secondary xylem) that increases stem diameter and transport capacity
xylem
- cells dead at maturity
- undergoes programmed cell death (PCD)
- loses cytoplasm and connecting cell walls forming long tubes
- heavily lignified, rigid
- way lignin is deposited varies
- provides some mechanical strength (primary source in small plants)
suction
- would not take H2O to tree tops
- maximum height of 10.6m = equilibrium point with a fixed vacuum taking into account strength of xylem
- after 10.6m gravity acts on water
mechanism of journey of water to treetops
- still debated
- main theory is cohesion tension model
- from models of synthetic tree (microfluidic system formed in a synthetic hydrogel)
- passive transport of water by capillary action
- adhesion and cohesion creates upward movement until balanced by gravity
- evaporation of water via the stomata from mesophyll cells in leaves (transpiration) creates tension (negative pressure) in the xylem)
- if H2O evaporates at top of column upward flow occurs to equilibrium height
cohesion
= attraction of like molecules
- water is polar
- 1 water molecules binds to 4 others (H bonds), strongly cohesive
- creates very strong surface tension
adhesion
= attraction of unlike molecules
- between H2O molecules and xylem wall from charge interaction
ABA and events in drought stressed plants
- drought stress simulated by witholding water
- stomata closure correlates with ABA levels (abscisic acid, plant hormone)
- modifying ABA response pathways could make plants more adaptable to drought as stomatal closure would happen more quickly, reducing water loss through transpiration
cuticle layer
- complex cutin biopolymer excreted
- waterproofs leaf surface, hydrophobic
- decreases evapotranspiration from upper epidermis
succulent leaves
- waxy leaves
- store water in fleshy leaves
- mucilaginous tissues within leaves can retain water through gel like properties
- reduces transpiration
- CAM photosynthesis
minimising leaf surface e.g. pine needles
- narrow elongated shape reduce stomatal pores and area of epidermal surface, decreasing water loss through evapotranspiration
- thick waxy cuticle coating, barrier against evapotranspiration and reflects sunlight (reduced thermal energy)
- sunken stomata in grooves creates microenvironment that reduces evapotranspiration
rolled leaves e.g. marram grass
- also has deep roots
- encloses internal space in rolled structure, les surface area exposed to direct sunlight and wind, creating microclimate
lithops ‘living stones’
- native to southern Africa
- camouflage, colour mimics geology, protects against herbivory
- grow mostly underground with just the leaf pair exposed to limit evapotranspiration
succulent stems e.g. cacti
- no leaves, stem photosynthesis decreases surface area and therefore evapotranspiration
- waxy cuticle
- sunken stomata into stem surface
- CAM photosynthesis, stomata open at night
- extensive shallow root system
- stem tissue adapted to store large amounts of water
- spines and hairs helps shade stem, collects dew and protects against herbivory