Perception Flashcards
What are sound waves?
waves of pressure changes in the air, caused by the vibrations of a source
What is Loudness?
- the percept associated with sound pressure level
- the amplitude of a sound pressure wave
- smaller amplitude = smaller loudness
- bigger amplitude = higher loudness
What is Loudness measured in and how can you measure it?
decibels (dB) - a physical measure of sound amplitude, or sound pressure level
measured by comparing two tones and deciding which one is louder
(measured in phones)
perceived loudness also varies with frequency
What is Pitch?
- the percept associated with sound frequency
- differences in pitch are due to changes in frequency of sound pressure waves
- high pitched tone = more waves / higher frequency of waves
- low pitched tone = less waves / lesser frequency of waves
What is Pitch measured in and why can it be considered to be a ‘metameric’ percept?
frequency is measured in (Hz) - a physical measure of sound oscillation rate
(1Hz = 1 oscillation per second)
metameric:
- you can get very different sounds giving you the same pitch
- e.g. the same note being played by a different musical instrument
Sounds are separated by an octave - same note but different frequency
- known as a ‘chroma’ which is basically pitch 2x
What is Timbre?
- the percept of the shape of sound waves
- has a negative definition - everything that is not loudness, pitch or spatial perception
- it is a multidimensional percept
- on instruments, you have the same notes but different frequencies
HARMONICS = integer multiples of the fundamental frequency - related to the relative amplitude of harmonics
- breaking down sound waves into its component processes
- adjusting the harmonics changes the timbre
Describe the physiology of the outer ear
Pinna and Concha
- gather sound energy and focus it via the auditory meatus onto the tympanic membrane (the ear drum)
- selectively filter sound frequencies to provide cues about the source’s elevation (where it comes from)
Describe the physiology of the middle ear
- tympanic membrane (ear drum) and the 3 ossicles; Mallues, Incus and Stapes
- role in impedance matching
- need to change the vibrations of the ‘sound’ so that the vibrations of the air match up the vibrations of the fluid in the cochlea
- done because air moves faster than liquid
- basically changes the amplitude - pressure boost up to 200x
Describe the physiology of the inner ear
- contains the cochlea
- vestibular canals which are important for balance
- where transduction happens; vibrations / pressure waves changed into neural signals
Describe the Cochlea and its functions
- coiled structure
- bisected from base to apex by two membranes:
1. basilar membrane
2. tectorial membrane - these contain the hair cells that do the transduction
- fluid (perilymph) that fills the chambers on each side
- vibrating because of the oval window (a membrane-covered opening at the base of the cochlea)
Describe the Organ of Corti
- the sensory organ which the equivalent of a retina
- runs along the whole of the basilar membrane
- contains hair cells with stereocillia transducers
- these convert the vibrations / motions of sound pressure waves into neural signals
What are the two different types of hair cells?
Inner and Outer hair cells
Describe Inner Hair cells
- the sensory receptors that send information to higher cerebral levels
- do the transduction - converting to neural signals
- 95% of the fibres in the auditory nerves
Describe the Outer Hair cells
- receive projections from upper cerebral levels
- roles in active filtering; modify the frequency of inner hair responses
- basically modify signals from the inner cells
Inner and Outer Hair Cells - Mechano-electrical Transduction
Vibrations:
- opens and closes ion channels
- by electrical depolarisation
- gives you action potentials to work with
- how mechanical vibrations are transduced into neural signals
What are the two important aspects in the Auditory Nerve?
Phase locking and Tonotopic Organisation
Describe Phase locking
need something firing at 20Hz - fire it at 20Hz in one go, keep sending these bursts of 20Hz?
- phase locking - in time firing
- one cue to sound frequency
- 260Hz at 260Hz - fire it at the right times
- each nerve fibre is tuned to a best frequency
What are the problems of Phase Locking and how can can we overcome this?
- nerve fibres can’t fire all the time - if something needs to be fired at 20Hz, may not be able to do that all the time?
- there are limits on transmitting
- can’t just do it along one fibre - so transmit multiple frequencies along multiple nerve fibres!
Describe the frequency decomposition along the basiliar membrane
Base = narrow and stiff; can’t vibrate very fast - low frequencies!
Apex = wide and flexible; can vibrate fast - high frequencies!
- different parts of the BM are ‘tuned’ to certain frequencies
What is Tonotopic organisation?
In the auditory cortex, different sounds are persevered in different areas
Pitch Perception: Place Model vs Rate Model
Place - argues that the position of excitation / vibration on the basiliar membrane determines how a frequency is encoded
Rate - encoding depends on the precise timing of individual spikes
The perceptual system appears to use a combination of the two techniques