Partitioning Flashcards
Energy balance
Sources - provide assimilates and energy; anything carbon based and anything photosynthetic
Sinks - use up assimilates; non photosynthetic parts; often edible
Energy losses due to inefficiencies
50% - out of photosynthetic spectrum
5% - reflected
6% - photochemical inefficiency (no conversion to sugars)
13% - thermodynamic limit
11% - carbohydrate synthesis (making enzymes etc)
6% - photorespiration
<5% of solar energy ends up as biomass
Photorespiration
Rubisco fixes oxygen instead of CO2 so wastes energy as useless product (2 phosphoglycoate) formed
Happens when CO2 levels are low
C3 vs C4 plants
C3 has more issues w photorespiration - horizontal layers of cells
C4 have reduced photoresp., increased WUE and photosynthetic efficiency
Concentrates carbon around rubsico and decreases oxygen
Spatial separation of processes
Circular cell arrangement
Photochemical inefficiency
Energy lost to heat; sometimes result of light: low light = more efficient as less energy lost as heat
Photosynthesis increases linearly w light intensity, then levels off when too high due to other limiting factors
Excess light dissapated as heat
Too much energy = free radical production and leaf damage
Max quantum yield - point where uses light most efficiently
Light compensation point - light required to overcome respiratory processes, but not enough for high photosynth than respiration
NPQ
Adaptation to deal w excess energy - photosynthetic proteins absorb excess e- energy and dissipate as heat
Slow to turn on and off to protect plant
Make NPQ more responsive to reduce energy loss - only want on when necessary
Xanthophyll cycle
Xanthophyll - pigment in plants; wide peak in blue only; good at stimulating conversion of excess e- energy into heat
Production of zeaxanthin from violaxanthin via antheraxanthin
Zeaxanthin - antioxidant to protect lipid membranes from free radicals
Partitioning
Make plant more efficient at putting energy where we want to eat
Products need a sink
Selectively breed plants depending on what we want to eat - different energy partitioning within plant
e.g. wheat - want high grain mass and fewer assimilates to roots and tillers
Sink strength can depend on stress
Harvest index
Fraction of crop mass that’s economically viable
Mass of product / Total above ground biomass
Transport - passive and active movement
Passive - e.g. osmosis: moving sugars along conc gradient; moves from high to low conc
Active - movement from low to high; want more sugar in sinks; requires energy for sugar transporters
Transportome
Move sugars and starch out of chloroplast and into cell and then other cell
Correct crossing of membranes via sugar transporters
High energy production
Control this pathway to control where sugars go for yields