Paleobiology Flashcards
Paleontology / Paleobiology definition
study of ancient life
science of the forms of life existing in former geologic periods- represented by their fossils
Geology
geo- earth, logia/logos- study of
the study of Earth
studies earth material- minerals and rocks
processes operating within and on earth
physical geology
examines origin and solution of earths continents, atmosphere, oceans, and life
historical geology
system
combination of related parts that interact
biosphere, atmosphere, lithosphere, hydrosphere
determined that what are now mountains used to be the sea by the fact that mountain rocks contained shells
Leonardo da Vinci 1452-1519
demonstrated that fossils represent remains of ancient animals
Niels Stenson / Nicholas Steno (1638 - 1686)
shark teeth
previously called tongue stone
glossopetrae (sharks teeth)
recognized that fossil record showed species appearances and extinctions
Robert Hooke (1635-1703)
Georges Cuvier (1769-1832)
Father of vertebrate palaeontology and catastrophism
catastrophism
history of earth can be explained by sudden catastrophic events
uniformitarianism
the assumption that the same natural laws and processes that operate in the universe now have always operated in the universe in the past
fossil
the remains or impression of a prehistoric organism preserved in petrified form or as a mold or cast in rock
Why fossils are important
simple fascination paleogeography paleoecology evolution biostratigraphy economics
compression
still contains parts of original organism
where can fossils be found
sediment, sedimentary rocks, metamorphic rocks, concretions/nodules
led to continental drift theories
paleogeography
paleogeography
environmental and physical restrictions to organism distribution
why fossils are important for evolution
fossils are the only direct record of the history of life
use of fossils in deducing succession and age relations, and dating sediments
biostratigraphy
index fossils must be
easily recognizable short stratigraphic range easily preserved (hard parts) worldwide/cosmopolitan distribution rapidly evolving abundant
an index fossil with a shorter stratigraphic range
gives more precise dating
economic fossils
chalk deposits ammonites for building structure dolomite mountains fossil fuels ( coal, oil, gas)
rudist
cretaceous reefs with high porosity and permeability, important for reservoirs and caps
process of fossilization
study of preservation
taphonomy
types of preservation
body fossils
molds and cast
ichnofossils
types of body fossils
unaltered remains
altered remains
unaltered remains happen
in unique environments where mechanical processes do not break down organisms
methods in which fossil organism may retain colour
freezing
types of unaltered remains preservation
Freezing
Drying/ desiccation
Amber/ tar/ wax/ asphalt
oldest fossil you could find from freezing
Quaternary
original materials for fossilization
CaCO3, Si, Apetite (calcium phosphate; teeth), chiton, cellulose, resistant organic substances (pollenin)
average ocean salinity
35ppt
photic zone
up to ~200m
freshwater salinity
<0.5ppt
brackish water salinity
5-30ppt
frozen fossils have
everything preserved - even internal organs
shore zone between high and low tide
littoral zone
below tide to edge of continental shelf, most diverse zone due to high nutrients and light
sublittoral zone
drifters, passive floaters
plankton
active swimmers
nekton
from continental shelf to abyssal plains
bathyal zone
benthic organisms on hard substrate
Epifaunal
benthic organisms on/in soft substrate
Infaunal (burrowing)
extract nutrients from sediment
deposit feeders
mobile marine organisms
vagile
immobile marine organisms
sessile
primary producers
cyanobacteria, algae
herbivores
gastropods
deposit feeders
gastropods, bivalves
suspension feeders
bivalves, crinoid
Amber fossils are mostly
insects
from tertiary
from baltic region
types of altered body fossils
permineralization
recrystallization
replacement
carbonization
another name for permineralization
petrification
permineralization
porous material fills with groundwater
original material not involved
heavier than original material
most common permineralization
wood
Si fills pores
recrystallization
hard parts revert to more stable minerals or larger crystals
example of recrystallization
aragonite—- calcite
in dissolution and replacement, hard parts are replaced by
calcite, silica, pyrite, iron
pyritization occurs in
anoxic environments
example of dissolution and replacement
ammonite turns from iridescent/ mother or pearl – pyrite
another name for carbonization
coalification
carbonization
only carbon remains, all other elements are removed, makes fossil appear black, organisms outline remains well preserved
common carbonization
organic rich shales, sandstones
plants (ferns), graptolites, fish
external / internal cavity
mould
filled in mould
cast
compression
2D shallow external molds that often display plant structures. retain original/chemically unaltered organic materials
impression
2D, no organic material, found in fine-grained sediment like clay or silt, commonly trace fossils, give insight into biological activity of organisms
tell more about organism behaviour than about the organism
trace fossils
types of trace fossils
tracks trails exogenic /endogenic trace fossil coprolite gastrolite
study of trace fossils
ichnology
vertebrate trace fossils
tracks
invertebrate trace fossils
trails
exogenic trace fossils
made on the surface of sediment (tracks)
endogenic trace fossils
made within the sediment (burrows)
fossilized poop
coprolite
fossilized digestive stones + stomach contents
gastrolites
only really distinct coprolite
sharks - swirly
coprolites & gastrolites tell
about diet not much else
set of trace fossils associated with a particular set of environmental conditions
ichnofacies
common / important ichnofacies
nereites, zoophycos, cruziana, skolithos, glossifungites, trypanites, teredolites
Nereites
deep marine ichnofacies
spiral, flower, sinuous, honeycomb
zoophycos
outer shelf - slope, low E muds, organic rich,
3D feeding trace (burrow, tube)
cruziana
shallow marine shelf- upper slope
unidirectional, continuous, crawling trace
skolithos
shallow, near shore vertical burrows (dwellings)
Glossifungites
in firm - not lithified- sediment (mud, silt), marine intertidal and shallow
plant root penetration, borings, burrows
Teredolites
borings in wood by bivalves
grazing traces are
very sinuous
resting fossil is
a depression in substrate
Trypanites
in hard substrate
predators (worms, bivalves, gastropods, barnacles) bore holes in corral, rock, shells
trace fossils characterized by behaviour
resting, dwelling, escape traces, moving, grazing, deposit feeding
pseudofossils
dendrites
concretion / nodules
solutioning
chemical / molecular fossils
chlorophyll, lipids
molecule characteristic of cyanobacteria found before earth was oxygenated
steranes evidence of eukaryotic life up to 1 billion years before they enter fossil records
dendrites
manganese, iron, tree like branching
how would bones be preserved
likely permineralization (porous)
how would teeth be preserved
likely recrystallization (solid)
preservation dependent on decay and mineralization
decay vs. mineralization, both must be minimum for soft parts to be conserved
mineralized muscle— tissue— chitin— cellulose— shells
steps of taphonomy
necrolysis
biostratinomy
diagenesis
necrolysis
decay and decomp., from death to right before break up
biostratinomy
mechanical processes, before burial
diagenesis
mechanical, physical, chemical, and biological changes after burial
biocenosis
life assemblage, interacting organisms in a habitat
thanatocenosis
assemblage of organisms brought together after death
taphonomic filter
more and more information is lost with every step, some organisms are lost throughout filter
most resiliant bones
lower jaw, skull
physical diagenesis
compaction, deformation
chemical diagenesis
dissolution, recrystallization, replacement
biological diagenesis
bacterial decomposition (early stages)
how settling of bivalves can tell about environment
if they land concave up and stay that way it must have been a low energy environment
elongate minerals/fossils settle in what direction
parallel to shore
important for determining compaction/deformation
cleavage- to not mistake as elongate fossils
missing from fossil record
organisms without hard parts organisms from unpreserved environments organisms rare or geographically restricted habitat info behaviour living morphology
how organisms with hard parts can be lost from record
if soft parts are all that distinguishes them from organisms with similar hard parts
lost habitat info
ex. elephant bones found in a lake- they don’t live there!
unpreserved environments
high latitudes (erosional environments)
communities lost from fossil record
only the biggest, strongest organisms are likely to survive, rare for young
material changed/moved due to erosion/reposition of sediment
reworking
reworking occurs by
lateral transport
storm/currents
bioturbation
examples of reworking
taxa ABC are eroded together- appear to have lived at same time but did not
2 halves of bivalve end up on different ends of beach
kelp beds end up on beach
sedimentation rate is so low that taxa ABC are deposited basically together
when a taxa disappears and then reappears although it never died
Lazarus taxa
lazarus taxa occurs due to
preservation issues
environmental changes
species endangerment
zombie effect
taxa ‘reappears’ after it has become extinct due to reworking
example of zombie effect
if an ammonite fossil surfaced now, but was reburied, and then resurfaced in the future
Elvis taxa
impersonating, morphologically similar organism to those before
Important for determining zombie effect
heat/colour alteration- sequence of colour change is irreversible, determines true age
how complete is the fossil record
> 97% is not preserved
preservation favours which environment
marine (and some aquatic)
‘motherload’ of fossils
Lagerstratten
two types of Lagerstratten
Konzentrat
Konservat
Concentration deposits
Konzentrat
concentrated but not best preservation
large number of fossils, low preservation of minute details
types of Konservat
Stagnation deposits
Obrution deposits
Conservation traps
Konservat deposits
best type of preservation, fine minute details are preserved
Stagnation deposits
stagnant water
supersaline
lagoonal
limits bacterial degredation (‘pickled’)
Conservation trap
Amber
tar pits
low decomposition
Obrution deposits
Organisms from normally not preserved environments are preserved
Burgess Shale- large chunk of shelf slid deeper
Biasis of fossil record
rapid burial anoxic/hypersaline condition no/minimal reworking or diagenesis tissue resistant to decay organisms in low energy environment marine organisms
grouping of objects or information based on similarities
classification
taxonomy
branch of biology concerned with the grouping and naming of organisms based on their similarities, chemical make up, similarities
by observing patterns we can
deduce factors that control organism distribution
six major kingdoms represented by fossil record
Plantae Fungi Animalia Protista (uni/multi cellular) Archaeobacteria Eubacteria
extremophiles
Archaebacteria- live in harsh environments (anaerobic, hyper saline, sulfurous hot spring)
cyanobacteria
filamentous, less tolerant than Archaebacteria
Protists
protozoans- dinoflagellates,diatom, foraminifera
Fungi act as
decomposers
parasites
kingdom Fungi includes
molds
mildew
mushrooms
yeast
plant evolution
green algae–spore plants (mosses)—vascular plants (ferns)— seed plants (gymnosperms, angiosperms)
living organisms are classified by
binomial system of nomenclature
Linnaean system
Species
can interbreed and produce viable offspring
basic unit of classification
organisms with structural, functional, developmental similarities
have unique two-part names
capitalized
genus name not species
italicized or underlined
genus and species
Classification hierarchy for plants
Kingdom Division Class Order Family Genus Species
classification hierarchy for animals
Kingdom Phylum Class Order Family Genus Species
mnemonic for classification hierarchy for animals
King Phillip Came Over From Great Spain
Paleontological species
based on similarities in morphology rather than genetic compatibility
Allopatric speciation
speciation that occurs when populations become isolated to an extent that prevents genetic interchange
sympatric speciation
one population slowly diverges from main population while still living in same area- don’t interbreed
Which speciation is easier to deal with in fossil record
Allopatric
Allopatric speciation can occur either
symmetrically or asymmetrically
Phyletic gradualism
theory that evolution is gradual at a ~constant rate
speciation occurs as slow gradual change
punctuated equilibrium
theory that periods of evolutionary equilibrium are interrupted by episodes of rapid evolutionary change
rapid change– relative stasis– rapid change–…
lineages show little evolution
punctuated equilibrium (stasis)
evolution takes place in lineages
phyletic gradualism
speciation is a side effect of evolution
phyletic gradualism
change in individual morphology during lifespan
ontogenesis
can compare species embryos
group of individuals of same species in an area
population
all populations of all species living in an area
community
a community and its abiotic environment
ecosystem
types of variation in organisms populations
ecophenotypic variation
taphonomic variability
sexual dimorphism
copes rule
ecophenotypic variation
change due to environment- nutrient, light, temperature
taphonomic variability
distortion after death (biocenosis vs. thenatocenosis)
biocenosis
describes the interacting organisms living together in a habitat
thanatocenosis
death assemblage
sexual dimorphism
phenotypic variation between males and females of species
cope’s rule
body size increases during evolution of a group of animals
types of skeletal growth
accretion
addition
molting
modification
Accretion skeletal growth
adding on discrete growth layers to the skeleton as organism grows
ex. corrals add layers every day, bivalves
recognizing sexual dimorphism
analogs to living organisms ex.antlers
geographic time/location- F/M wouldn’t be thousands of years apart in record
Ratio ~50/50 M/F
similarity at early life stages
Addition skeletal growth
adding discrete new parts which grow very little after formed
ammonite, foraminifera (chambers)
Modification skeletal growth
continuous remodeling and adding to existing skeletal elements- mammals
Molting skeletal growth
shedding of exoskeleton- trilobite, crabs
Steno’s three laws for sedimentary rocks
Principle of superposition
Principle of original horizontality
Principle of original lateral continuity
Physical principles of relative age
Principle of superposition, original horizontality, original continuity, cross cutting relations, inclusion, recorded history, unconformities, fossil succession, fossil correlation
principle of superposition
youngest strata is on top
principle of original horizontality
sediment is deposited in horizontal layers due to gravity
principle of original lateral continuity
continuity is preserved from one environment to the next
principle of recorded history
using known facts to date (ex. volcano eruption)
1986 peak in Cs level
chernobyl
principle of unconformities
surfaces of erosion or non-deposition (hiatus) include significant amounts of geologic time
principle of fossil succession
oldest fossils in a series of sedimentary rock layers will be found in the lowest layer
principle of fossil correlation
similar assemblages of fossils are of similar age and therefore the strata containing them are of similar ages
chronology of events in Earth history established on basis of obtaining ages of past events
Geological Time Scale
chronometers
radioactive elements
can measure up to ~7 half lives
if radioactive element has long half life
not able to measure daughter products in young materials
if radioactive elements have short half lives
cannot measure daughter product beyond certain time
elements for absolute dating
Pb210
Cs137
Pb210
good for <120year old sediment
half life 22.3 years
Eons
Phanerozoic
Proterozoic
Archean
Hadean
Phanerozoic
visible life
Proterozoic
early life
Archean
ancient life
Hadean
greek mythological hell (Hades)
~90% of earth history
Precambrian: Proterozoic + Archaen + Hadean
Era
precambrian
paleozoic
mesozoic
cenozoic
Period mnemonic
camels often sit down carefully perhaps their joints creek TQ
geological time- periods
cambrian, ordovician, silurian, devonian, carboniferous, permian, triassic, jurassic, cretaceous, tertiary, quaternary
we no classify carboniferous as
upper- Pennsylvanian
lower- Mississippian
biozone
body of rock whose lower and upper boundaries are based on the range of one or more taxa
at well established geological time boundaries there are
golden spikes
what defines biozone boundaries
large extinctions
no fossil records or rocks of
Hadean
epoch time unit
early/middle/late
epoch time-rock unit
lower/middle/upper
Lower Devonian rocks represent Early Devonian time
history of Earth
4.6 billion years
time of Hadean
4.6-3.8bya
time of Archaen
3.8-2.5bya
time of Proterozoic
2.5bya
time of Cambrian
542mya
time of Silurian
444mya
time of carboniferous
360-250mya
time of mesozoic
250-65mya
time of tertiary
65-2.6mya
time of quaternary
2.6mya-present
time of cenozoic
65mya-present
by end of Archean
earth had an atmosphere, greenhouse gases, plate tectonics, life
evidence of Precambrian life
morphological fossils (black chert)
stromatolites
chemical fossils
chemical fossils
C12/C13 ratios, organisms preferentially take up lighter C12
Pristane/phytane evidence of photosynthesis
prokaryote size
~10µm
how long was all life on earth bacterial
~2bya
beginnings of life on earth
Isua formation of greenland
Warrawoona group
fig tree formation
gunflint cherts
Isua Formation of Greenland
3.85bya, oldest altered sedimentary rocks, geochemical indicators, carbon isotopes
Warrawoona Group
Apex chert, Western Australia, 3.5bya, stromatolites, six types of filaments
Fig Tree Formation
3.4bya, South Africa, cyanobacteria filaments associated with stromatolites, light carbon isotope ratios, pristine and phytane in cherts
Gunflint cherts
Canada, 2.1bya, stromatolites, black cherts, chemical fossils, diverse prokaryotes, numerous bacteria types
oncolite
similar to stromatolites, instead of forming columns, they form spherical structures, often form around central nucleus, shell fragment, and calcium carbonate structure is deposited by encrusting microbes
most likely origin of life
hydrothermal vents (chemoautotrophs)
first life
early archaen
why hydrothermal vents are a likely source of life
abundant energy/mineral supply
ocean protects from UV
many prokaryotes live near vents
needed for life to occur
cellular structure
metabolic assimilation of energy
reproduction
heredity
origin of life theories
creation
extra terrestrial
spontaneous generation
inorganic model
inorganic model
genetic material evolved first in association with clay minerals and organic compounds were involved only later
hydrogen oxidation
2H2 + O2 —- 2H2O + energy
sulfur reduction
S + H2 – H2S + energy
methane production
CO2 + 4H2 – CH4 + 2H2O + energy
first photosynthesizer
cyanobacteria
kingdoms in Archean
archeobacteria
eubacteria
free oxygen (Tertiary atmosphere)
2.5-26mya
BIFs
interbedded chert and iron rich minerals (Fe sulphides, Fe carbonates)
need low level oxygen (precambrian)
BIFs first appear
3.8BYA (more common in proterozoic)
BIFs rare after
1.9BYA
indications of higher levels of oxygen
stromatolites
eukaryotes
oxygen ‘sinks’ start to fill up and O2 can accumulate in atmosphere
~2BYA
Proterozoic eon life
2.5-0.5BYA
evolution of complex eukaryotes
first multicellular life
evolution of sexual reproduction
origin of eukaryotes
2-1.8BYA
origin of sexual reproduction
1.1BYA
origin of multicellular life
~0.7BYA
evidence of Proterozoic evolution
gunflint chert
bitter springs chert
Rodinia
supercontinent ‘motherland’
1.2BYA-600MYA
Breakup of Rodinia
snowball/ slush ball earth
extinction- glaciation prevents light
Ediacara fauna
also Vendian (soft-bodied animals) developed movement, symmetry very thin, up to 1m some anoxic species- maybe symbiosis possibly leathery no predation, filtering/grazing
Ediacaran phytoplankton
Acritarchs- main primary producer after cyanobacteria
organic walled, unknown affinity
resistant to dissolution
Cloudina sp.
first skeletal fossil (Ediacaran)
CaCo3, CaPO4, tube dwelling worm
Pre-late Cambrian
Tommotian Fauna
small shelly fossils CaPO4 first evidence of predation (protective pieces) evidence of competition (grew taller) 1-5mm
Cambria diversification
continents split up, new ecological niches, sea level changes, expanded cont. shelf, warm water, transgression all phyla (except bryozoa) appear VERY rapidly
transgression
sea level rises- shoreline moves inland- mud deposited directly on old beach sand
cambrian life
many more shelled species
decreased stromatolite abundance
all phyla (except bryozoa) appear VERY rapidly
most major invertebrate classes
hard part advantages
protection from UV- can move to shallow water
prevents drying in intertidal pool
predation protection
support
hard part disadvantage
energy of molting
Archaeocyathid
sponges, appear in Tommotian (early Cambrian)
extinct by mid Cambrian
1-3cm diameter, 15cm tall
cone-in-cone structure, pores, central cavity
calcareous skeleton
benthic, sessile, colonial or individual
excellent indicator of early Cambrian
Archaeocyaths- only marine, benthic, sessile, passive filter feeders in 20-100m water depth of tropic carbonate shelves
Kingdom, phylum, subphylum of trilobite
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Trilobitomorpha
Class: Trilobita
Trilobite time frame
only in Paleozoic extremely common in Cambrian and Ordovician early Cambrian- end of Permian 8/9 orders appeared in Cambrian 15,000 species
Trilobite exoskeleton
calcite
main index fossil of cambrian
trilobite
Major body parts of trilobite
Cephalon, Thorax, Pygidium
Pleural lobes, axial lobe
evolutionary trends of trilobites
greatly elongated to transverse (widened)
thoracic segments increased (60) or reduced (2)
reduction/loss of eyes
Effacement
adaptation related to a burrowing lifestyle
smoothing out of lines for burrowing (or streamlined swimming)
Spinosity
defensive/stabilizing adaptation
can tell modality from direction spines stick out
pelagic trilobite morphologies
extremely large eyes
streamlined bodies
olenimorph
forms associated with low O2, high sulfur benthic habitat
thin exoskeleton, increased thoracic segments, wide flat body, symbiotic behaviour
why wide flat body in olenimorph
larger surface area : water interface for material exchange
environment changes for trilobites mostly related to
temperature changes- largest effect in shallow water
Burgess Shale
mid Cambrian
quick burial, anoxic conditions
soft parts preserved
~93 soft bodied organisms
one of the first large predators (seen in Burgess shale)
Anomalocaris canadensis (sea horse kinda shape)
can tell predator from distinct bite marks in trilobites
up to 60cm
earliest chordate
Pikaia
~5cm, nodochord, zigzag muscles attached to nodochord
Burgess shale organism that was incorrectly reconstructed (upside down)
Hallucigenia
~1cm long, protective spikes on back
why explosion in Cambrian
few predators increased atmospheric oxygen sea level rise (many habitats) evolution of hard skeleton higher nutrient levels
parts of evolutionary faunas graph
mud grabbers–stationary filter feeders–mobile filter feeders–soft bodied organisms
1-5 large extinction
plateau in cambrian- max # of species that can live on mud substrate, need grow taller evolution
majority of species diversification at end of Cambrian
cambrian fauna
trilobite fauna
mainly trilobites, also echinoderms, brachiopods
paleozoic fauna
brachiopod fauna
brachiopods, bryozoans, graptolites, cephalopods, crinoids
Ordovician changes
485-443mya north of tropics open ocean most land masses southern supercontinent, Gondwana O2 at modern levels flooding of continents
Ordovician radiation
150 families in Cambrian
400 families in Ordovician
sediment changes from Cambrian-ordovician
Cambrian flat planar beds
Ordovician bioturbated, by upper Ordovician can’t tell bedding planes
Tiering
increase height and depth in substrate
Conodonts
worm like forms with teeth, CaPO4, change colour in irreversible way, CAI, teeth not attached to jaw
marine, free swimming, pelagic and nektobenthic predators, carnivores or scavengers
conodont phylum
EITHER Hemichordata or Chordata (uncertain)
conodont importance
extinct in Triassic, good for dating between triassic/jurassic, if find conodonts KNOW its not Jurassic
conodont morphotypes
coniform
ramiform
pectiniform