Palaeobiology Flashcards
Definition: The branch of palaeontology that deals with the processes of fossilization
Taphonomy
Definition: The physical and chemical processes that affect sedimentary materials after deposition
Diagenesis
To be a fossil, an organism must:
1. Die and avoid destruction
2. Be in an environment where they can be buried
3. Be fossilized
4. Avoid d—— or m——- processes
5. Avoid e—–
6. Become e—— and discovered
diagenetic, metamorphic,
erosion, exposed
Fossils can have a n—— cast (rarer) or an i—– cast
natural, internal
Why is the fossil record incomplete?
1. S—— only accumulates on small area of Earth
2. Only small amount of organisms fossilized
3. Most d——— with the rock they are in
4. Only a t— fraction found & studied
sediment, destroyed, tiny
Why is the fossil record biased?
1. Certain o——- parts preserve better
2. Certain e——— preserve better
3. O—- rock more likely to be destoryed
4. C——— bias ie. First-world countried
Organism(s), environments, older, collector
Which kingdom leaves these parts fossilized?
Stromatolites and certain bacterial sheathes
Bacteria
Which kingdom leaves these parts fossilized? Those with an exo or endoskeleton.
Protists (single-celled living things)
Which kingdom leaves these parts fossilized? Woody tissues made of lignin. Cuticles made of Cutan. Spores known as sporopollenin.
Plants
Which kingdom leaves these parts fossilized? Chitnous spores and hyphae. Unfortunately these are often similar across species and the ‘useful’ part is lost.
Fungi
Which kingdom leaves these parts fossilized? Those with recalcitrant exo and endoskeletons
Animals
In which environments do sediments accumulate?
- C——– S—-.
- Rarely in deep sea r– c— from the atmosphere
- OCCASIONALLY in inland f—– in the mountains
continental shelves, red clay, faults
Large-scale biases of fossil distribution:
1. S–Level changes
2. Continental c——-. More s—- when continents SPLIT
3. Atmospheric composition. More o—— = more composition
sea, configuration, shelf, oxygen
Definition: It is probable that patterns in the fossil record reflect both biological and geological signals. We must therefore understand the geological signals
The common cause hypothesis
How to interpret the fossil record;
- Plot s—– and t——- distribution of fossils
- Consider g— ranges where fossilization didn’t occur
- Consider probabilities of r— extensions
- Analyze volume of r— depositied per time slice and the e——- area of rock per time slice
spatial, temporal, ghost, range, rock, exposed
The fossil record is GOOD to:
- Give us a t—f—- for evolution
- Tells us what organisms e——- from and what wetn e—– / why it did
time-frame, evolved, extinct
Dinosaurs are GOOD to study due to having b—- and t—- but BAD as a lot were t——–
bones, teeth, terrestrial
Dinosaurs first evolved in the T—— and lived through the Jurassic and the C———
Triassic, Cretaceous
What is a Fossil-Lagerstatten?
a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues.
What are the two types of fossil-lagerstatten?
1. ———- deposits such as bone beds, mass kills and strand deposits
2. ——- deposits where preservation is of unusual quality
Concentration (where fossils occur in unusual concentrations), Conservation
What can create a bone bed?
A landslide
What can create a mass kill?
A pond drying out or something
What creates a strand line deposit?
The tide
What are the 3 basic types of conservation deposits?
1. Preservation of o—— which aren’t usuually preserved.
2. Preservation of p—- of organisms which aren’t usually preserved (skin)
3. Organisms preserved in unusually a———- or unusual configurations (ichthysaur giving birth)
organisms, parts, articulation
What is this a definition of?
The reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains.
(a) bioturbator
Give 2 major controls for exceptional preservation. Why may they be no bioturbadors?
- No scavanegers and bioturbators, perhaps due to anoxia, rapid burial or elevated salinity
- Unusual chemical environment permitting exceptional preservation. May be due to a catastrophic event or bacteria producing certain chemicals
During deposition, the formation of ——- and/or ——— mats may help preservationq
Nodules, microbial. Stabilized sediments may set up geochemical gradients that promote early mineralizations.
Give the 6 conservation traps that we learnt about. What is a hot silicious spring?
- Amber
- Ice- Ice only goes back about 1 million years.
- Tar Pits - La Brea tarpits in LA from the Pleistocene.
- Hot Silicious springs- Water is so hot it dissolves silicate, This silica traps all plants around it.l
- Tufa - In a limestone environment. Slightly acidic rain dissolves limestone and forms stalagmites.
- Ash falls - pyroclastic flows. Very fast. This happened in pompeii
What is preserved well in ice?
Meat. Can show forms that bones alone cannot
What is preserved well in amber?
Insects and small creatures
What is preserved well in tar pits?
Bones.
What is preserved well in hot silicious springs?
Single-cells can be!
What is preserved in Tufa?
Birds nests and things!! in caves
The process of exceptional preservation.
1. Burial
2. W— pushed out by pressure
3. M——- left behind.
4. Organism can d—– in the water. Gap filled with precipitated minerals
Water, minerals, dissolve
In prservation, occasionally loads of m—— is precipitated out to form a n—–
mineral, nodule
Tissues display a spectrum of resistance to decay from re——– tissues to l—— (more soft) tissues such as muscle
recalcitrant, labile
Decay proline tissues are only preserved when replicated fast by authigenic minerals which:
1. Precipitate round the tissues and form a c—
2. Precipitate on and within the tissues infilling and potentially preserving actual c—–
cast, cells
What is the best mineral for preservation? Where does it come from?
Apatite. It can preserve muscle. Is released from phosphate in an organism when it dies.
In the past, p——– levels may have been allowed to build up due to absence of life forms using them, allowing organisms to be perfectly preserved in them.
Phosphate
What are authigenic materials?
Those which have not moved from the position in which they formed.
Explain how coal allowed for exceptional preservation and why this happened more in warm wet environments. What does coal normally preserve? How could it form in cold environments?
Coal minerals can also allow for exceptional preservation, we just don’t aren’t sure how this happened in cold environments, as coal often formed from environments where decomposition was higher, which were once warm and humid where there were lush plants growing. Coal mainly preserves plants and their pollen / spores / cuticles Also more associated with swampy conditions as these would form the environment necessary for its formation.
Could ALSO form in cold environments in peat bogs, permafrost, glaciers or cold water swamps or wetlands
Exceptional preservation can also result from:
- — —– (fools gold)
- Other —– sulphides
- —– in hot springs, especially good for wood
- —— (can form coal balls - a lump of petrified plant matter)
iron pyrite, metal, silica, calcite
Give 3 ways in which Fossil-Lagerstatten confuse us
- False biodiversity peaks
- Preferentially occur in certain time periods
- Certain environments overrepresented
Example of exceptional preservation: The —– (Cenozoic Period, Germany)
- Created by v—- activity.
- P——– different parts in different organisms
- C– killed things
Messel, Volcanic, preserved, C02
Example of exceptional preservatin: ——– state (Devonian period, Germany)
- Iron pyrite and f—- black mud preserved s– organisms and not just their exoskeletons
Hunsruck, fetid, sea
Case study of the —– Biota (C———–, China)
- Formed due to volcanic l—- forming a– beds. Ash k——- out organisms leaving them at the base of the lake. F— grain could preserve in detail.
Preserved organisms AS THEY SWAM - behaviour. Told us REAL f—– colours.
Jehol, cretaceous, lakes, ash, knocking, fine, feather
Studies by Zhou
Hypsilophodontidae and i——– were a very diverse group of bipedal dinosaurs includeing the h——- (duck-billed) dinosaurs. Were quite intelligent.
Iguanodontia, hadrosaurs
The s——- included the odontosauruses and were very big and surprisingly diverse. Form a sister group to the t——–
sauropods, theropods
What were the theropods?
Bipedal carnivorous dinosaurs. Varied massively in size and were intelligent so as to be dinosaurs.
Birds were the sister group of d———–
dromaeosaurids.These had bigger brains to navigate 3D environments Birds are clearly theropods, a monophyletic group.
Time periods were s——— into different stages
subdivided
Iron minerals a—- towards the poles. Rocks can be analysed and relative m——— content can show where rocks were in relation to the poles, helping to map intial continental configuration.
align, magnesium
Using the position of s———- rocks today can indicate the rock was in a particular e——– initially due to the processes causing that sediment to be deposited there
sedimentary, environment
Attaining the original positions of rocks must account for s– level changes
sea
T/F To an extent the religious people and assumption that geological processes happened back then were the same was right
True (Was not noahs arc and environment HAD changed since then however. People gradually realized this)
Explain the 5 ways in which we can reconstruct past environments?
1. Examining the nature of sediments
2. Using the nature of a rock’s fossil content
3. The distrubution of climactically sensitive sediments and fossils
4. Environmentally discriminating isotopes.
5. Modelling
- Do this by relating it to today’s landscapes
- Using species present as a kind of historical ‘bioindicator’.
- Certain sediments only form in certain environments, like coal, bauxite, evaporite etc.
- Different composition of certain isotopes like boron form in different conditions of pH / temperature etc.
- Study impact of climate today and applying it to past variables. Loses efficiency the further back in time that you travel.
What does the ‘pull of the recent’ describe?
The fact it is easier to get data from EXPOSED sites which will tend to be more RECENTLY formed rock
Dinosaurs first appeared when the c——- were together thus there was less b———- variation than later on
continents, biogeoraphical
Why is oxygen concentration harder to calculate from past records?
Oxygen levels. Oxygen is more reactive so doesn’t leave direct signatures for us to read unlike C02 trapped in ice cores and such. Oxygen also constantly cycled round environment, unlike C02 which can end up stored. Must used proxies like concentrations of certain isotopes to infer it’s presence
Climatically s——– indicators used to guess what world dinosaurs lived in
sensitive
Climactically sensitive indicators tell us that the polar dinosaurs lived in a c— and t——— world.
Cold, temperate
What do LAGs tell us about the polar dinosaurs
Lines of Arrested Growth tell us that the polar dinosaurs did NOT hibernate
What are the ‘strange groups’ found across the construction of the dinosaur phylogenies
Charles’ answer: Basically these animals are anatomically strange (with weird primitive characters and missing some derived characters) and therefore fall out as basal within the phylogenies. I believe they form paraphyletic groups, maybee reflecting gaps in the fossil record and our own current understanding
How do we reconstruct an animal from what we find fossilized?
Make inferences based on the nearest living relatives . Also flesh it up using exceptional preservation cases and looking at muscle attachement sites on bone.
We can reconstruct extinct species capabilities using b——– and p——. We can also use 3D computer programmes to reconstruct them. T——- tell us about their height, speed andf stride-length
biomechanics, physics, trackways
What kind of dinosaur was the fastest? Which was the slowest?
The small bipedal theropods were the fastes wheras the quadrapedal stegs and anklyosaurs were the slowest.
How do we reconstruct an extinct species intelligence?
Using brain:body ratios. Fun fact - birds got more clever to navegate 3D space when flying.
Senses were easy / hard to reconstruct. Can compare with the modern reptile brain to do this
Hard
Ways to reconstruct a species diet:
1. T— form
2. J– adaptations - reptile herbivores must d——- them to do so.
3. Co-o——–
4. C——– ! Hard to link producer to content
5. G– contents
6. Fo——- finds - velociraptor / protoceratops death lock
- teeth
- jaw
- co-occurance
- coprolites
- gut
- Fortuitous
Ways to reconstruct behaviour:
1. E—. Ossification level of offspring in them, buried or not. Less ossification may show higher levels of parental care and investment
2. T——- - parallel? Followed by predators?
3. S—— displays - frills / pachycephalosaurs.
eggs, trackways, sexual
What limited the size of the biggest dinosaurs?
Pumping blood, drinking water, sinking in mud.
Here is some evidence for dinosaur thermoregulatory physiology. Google the ones you don’t understand.
There are 10 in total.
- Stance and activity levels
- Adaptations for processing high volumes of food
- Haemodynamics - Structure of heart
- Brain size - Bigger brain = more energy needed.
- Nose morphology - may be adapted to heat up air as it goes in.
- Bone Histology and Growth rates - can indicate metabolic rates. More blood vessels in bone = higher metabolism
- Predator - prey ratios - MORE prey would be needed to support warm-blooded animals
- Palaeogeography - where did they live? d
- Core and peripheral temperatures. Look at isotope composition of peripheral versus internal bones to indicate heat distribution in their skeletons.
- Presence of feathers?
Give some theories of what the plates on stegosaurus could have evolved for
Thermoregulation
Flapping down for defence.
Could have been spines originally then evolved into plates.
T/F Darwin believed that we could see evolutionary patterns in the record
FALSE he thought we could not but this has been disproved
Give the Evolutionary Patterns based on their description here:
1. - Variety across all of life, different kingdoms & clades.
2. Describes how morphologically disparate (variable) a certain group is.
- Describes what originated from what and at what rate species originate - punctuated equilibrium. Is this pattern across all groups at the same time or across time?
- Is there a background rate of extinctions? Do extinction rates cluster across a certain time, and is this across all groups or not?Not all groups DO follow these same patterns.
There is a need to differentiate between background extinction, pulses of extinction (extinction events) and mass extinctions. - How long do taxa live between extinction events? Chances of going extinct are believed to be INDEPENDENT of how old you are → Similar to that essay you did :). Age does NOT really correlate with extinction, maybe more to do with red queen hypothesis or such.
- Stable or in bursts? Can be related to morphology, genomics or taxa.. Can see if in phylogenies species accumulate slowly or appear to ‘bunch up’ in certain places throughout time.
- Plotting where fossils occurred when they were alive. Can plot occurrences of fossils across a phylogeny and see how these species spread across the plane
- Biodiversity
- Morphological disparity
- Origination Patterns
- Extinction Patterns
- Taxonomic Duration
- Rates of evolution
- Fossils & Biogeographic patterns
Give the Big Mass Extinction based on their description:
1. Short glaciation event, killed lots of tropical shallow reef biota
2. - Seems to be pulsed due to probably aquatic anoxia, related to the spread of forests, possibly. Vegetation caused algal blooms then anoxia in oceans
3 - Related to atmospheric pollution & heating due to the emplacement of Siberian Traps Large Igneous Provinces (LIPs). This lava released loads of gas and caused lots of acid rain. The sea went above a certain temperature and metabolism broke down in animals, especially those living across the equator
4 - Another LIP in the Central Atlantic. Poisons the atmosphere.
5 - Bolide impact (the meteor) caused a nuclear impact
- End Ordivician
- End Devonian
- End Permian
- End Triassic
- KT
Define this: the problem that due to the incomplete nature of the fossil record, it is extremely uhnlikely to find the first or last organism of any given taxa in a fossil
The Signor Lipps Effect
It is difficult to study mass extinctions as you need long c———- stratigraphic sections of MANY h—— before and after the event
continuous, habitats
What is this: refers to the fact that to an extent, which organisms survived a mass extinction could be largely stochastic
Gould’s contingency
Recovery after a mass extinction can be rapid due to a—— r—— or protracted due to too few taxa remaining to d——
adaptive radiation, diversify
What are Lazarus taxa?
Taxa which disappear after an extinction event but then arise again in a small population (named after a saint that Jesus resurrected)
What are Elvis taxa?
(Think Elvis impersonators)
Taxa which look like something which evolved before but did in fact evolve independently
What are Lilliput taxa?
A small / dwarf taxa which survives a mass extinction (Lilliput is from Gulliver’s travels)
Phyletic gradualism is now less believed than punctuated equilibrium. Supported by Gould and Niles 1993. When does punctuated equilibrium evolve?
When a small isolated population quickly evolves to be very different species which diversify. Happens in bursts. Peripatric speciation
Define these terms used to talk about punctuated equilibrium
1. Species sorting
2. Species selection
3. Habitat tracking
- Fledgling species are more likely to survive if they are ecologically different from their parent species (less competition) – this idea talks about niche partitioning essentially
- The differential rates of appearances and extinction of species within lineages –. The TURNOVER of species
- Ecological communities follow habitats suitable for them during environmental change.
Evolution is tirggrered by BOTH the red queen andf the court jester. Who is the court jester?
Just the environment changing rapidly.
What are the different tiers of evolution?
First Tier-
Second Tier -
Third Tier -
- More species level. Evolutionary events in the ecological moment on the day to day.
- Punctuated events and trends withing lineages over GEOLOGICAL time caused by isolate between pops.
3, Mass extinctions, wiping out whole groups and allowing new ones to arise.
Which species was evidence for the origin of birds?
Archaeopteryx
Birds are highly d—– dinosaurs evolving specifically from the t——–
derived, theropods.
Define paleodiversity
Considers how biodiversity has varied in the past
Which are the two ways in which the taxic approach can study paleobiodiversity?
- Can examine the rate of discovery curves. When the curve plateaus we can infer all species have been discovered (like in SARs)
- Can extrapolate from an intensive local sample (eg. one tree) and then estimatr how many undiscovered species there are
Try and label some hings which may be hard to recognize in the fossil record, for example the presence of diseased individuals.
- Species, ontogenetic stages, sexual dimorphism, ecophenotypes
It is hard to know if Linnean Hierarchies (Kingdom phylum class etc.) are e———- across different groups of organisms
equivalent
To assess paleobiodiversity, at a prescribed interval in time, you can measure:
1. M——— diversity
2. The number of t— –> Works best if they are well known, well represented and well preserved. Can use higher taxa as proxies for taxa number
morphological, taxa
What percent of the marine organisms which have ever lived have we probably described?
2-4 %. This is the same estimate for the % of organisms which ever lived which live in todays world.
What does the PDB stand for?
Paleo-database. It shows the presence of certain taxa through time.
What are the 3 models for the ways biodiversity can increase?
1. A—– caused by irregular extinction
2. L—— with a slow start, exponential growth then a plateau
3. E———. In a bifurcated model where rate starts slow then keeps rapidly increasing.
Different taxa show different models of these
additive, logistic, exponential
Why may life be more diverse on land?
Land it more heterogeneous. Mainly due to insects and soil microbes but we also do NOT know much about sea microbes
Explain these 2 explanations for patterns of biodiversity
1. The Equilibirum Model
2. The Expansion Model
- There is a ‘ceiling’ on biodiversity due to diversity damping factors (like competition) or limiting equilibirum factors (like densitiy dependence but for species)
- There is NO biodiversity ceiling. Seems unlikely.
There is much controversy as to if the e– C——– demise was real or not. May be down to rock available to study. Discusses across many papers on dinosaur diversification
end-cretaceous
It is hard to get DNA from fossils, as the instant something dies it becomes infested with external & internal m——- which c——— it, and DNA breaks down VERY quickly in the presence of w—- or a–
microbes, contaminates, water, air
We have only got SOME dna preserved from m——- in ice, egyptian mummies, and g—- sloths.
mammoths, ground
Rapid technologicalk adances jave allowed us to sequence DNA of many e—– species, like c— bears, Irish e–, N———, mammoths and more.
We have even obtained T.rex p——- sequences from bone-derived collagen.
extinct, cave, elk, neanderthals