Pages 492-499 Flashcards
Speciation begins when
gene flow between populations is reduced or eliminated, causing genetic isolation.
Genetic isolation
happens routinely when populations become geographically separated. Happens in one of two ways: dispersal or vicariance, the physical splitting of a habitat.
allopatry
populations that are geographically separated.
allopatric speciation
speciation that begins with geographic isolation.
biogeography
the study of how species and populations are distributed geographically.
Dispersal and colonization, followed by genetic drift and natural selection, is thought to be responsible for speciation
in the Galapágos finches and many other island groups.
sister species
species that are each other’s closest relative.
Geographic isolation of populations via dispersal or vicariance produces genetic isolation due to
the interruption of gene flow – the first requirement of speciation. When genetic isolation is accompanied by genetic divergence due to mutation, selection, and genetic drift, speciation results.
sympatry
when populations or species live in the same geographic area, or at least close enough to one another to make interbreeding possible.
sympatric speciation
speciation that occurs even though populations live within the same geographical area.
Two types of events can initiate the process of sympatric speciation:
- External events, such as disruptive selection for extreme phenotypes based on different ecological niches.
- Internal events, such as chromosomal mutations.
niche
describes the range of ecological resources that a species can use and the range of conditions that it can tolerate.
Even though sympatric populations are not geographically isolated, they may become
reproductively isolated by adapting to different ecological niches via disruptive selection.
Natural selection and genetic drift are usually responsible for amplifying the effects of
mutation.
polyploidy
occurs when an error in meiosis or mitosis results in doubling of the chromosome number – a massive mutation.
Autopolyploid (“same-many-form”)
individuals are produced when a mutation results in doubling of chromosome number and chromosomes all come from the same species. Less common than allopolyploidy.
Allopolyploid (“different-many-form”)
individuals are created when parents that belong to different species mate and produce an offspring with two different sets of chromosomes.
Polyploidy individuals are reproductively isolated from the original diploid population
and thus evolutionarily independent, because breeding between diploids and tetraploids generally results in sterile offspring.
Offspring is sterile if it has chromosomes that do not
pair normally during meiosis. Although, if a mutation occurs during mitosis that doubles the chromosome number in this individual before meiosis, then each chromosome gains a homolog and meiosis can proceed normally.
Why polyploids have been so successful:
- Polyploids have higher levels of heterozygosity than their diploid relatives.
- Polyploids can tolerate higher levels of self-fertilization because they are not as affected by inbreeding depression as their diploid relatives.
- Genes on duplicated chromosomes can diverge independently, increasing genetic variation in a population.
- Speciation by polyploidization is virtually instantaneous.