Molluscs Flashcards
Where are the molluscs found and how many species are there?
They are found in the sea, in freshwater and on land with ~100k spp.
What are Molluscs?
A major radiation of the metazoans. The name means soft bodied.
Where did molluscs originate?
They have a marine origin and are an ancient phylum which can be traced back to the precambrian. The earliest molluscs are the Helliconellida (540Mya)
What is the basic Molluscan body plan?
+A shell, usually CaCO3
+A mantle - holds everything together.
+A circum-oesophageal ring - the ‘brain.’
+Mouth, containing radula.
+Digestive gland, where enzymes are collected.
+An open circulation system.
+An excretory organ where nitrogenous waste/ammonia is removed
+Gills
What is the circulatory system of a mollusc like?
The heart contracts and pumps haemolymph to areas that require high blood pressure, like the excretory system.
What is the size range in the mollusca like?
1mm->9m.
What is molluscan taxonomy like?
The mollusca are polyphyletic - all groups do NOT share an immediate common ancestor.
What are the main classes, subclasses and orders contained within the Mollusca?
Bivalvia (scallops, clams, mussels etc.)
Monoplacophora (limpet-like ‘living fossils’)
Gastropoda (snails, slugs, limpets, sea hares)
Cephalapoda (squids, octopuses, nautilus, ammonites)
Scaphopoda (tusk shells)
What are the Aculifera?
A group of organisms:
Aplacophora (spicule covered worm like animals)
Polyplacophora (chitons)
Commonality of ancestry so far back that they have little in common. Argued they could have their own phyla.
What are the Aplacophora?
Aculiferan molluscs, classified as such due to their radula and other molluscan characteristics in internal anatomy.
They have spicules in the outer layer.
What are the two classes of Aplacophora?
The Chaetodermomorpha: wormlike burrowers with spicule-reinforced outer layer.
The Neomeniomorpha: they have a creeping foot-like ridge. They are unsegmented.
What are the Polyplacophora?
Name means ‘many plates.’
Chitons are an ancient exclusively marine group.
Shell consists of eight interlocking hinged plates, with a fringe of spines or spicules.
What are the function of the spines or spicules in the Polyplacophora?
The plates deter predators attacking from the top, so the spines or spicules are there to deter predators from attacking from the side.
What are the Monoplacophora?
Shell consists of single cone of material. Discovered in 1952. 10spp now known. They are deep sea.
Paired arrangements of gills and gonads are replicated, not evidence of vestigial segmentation.
What are the Gastropoda?
Name means ‘stomach foot.’
Soft tissues covered by a single shell, usually tubular and coiled on itself. Where external shell has been lost, vestigial structures are found in the soft tissues.
What is gastropod torsion?
Within the soft tissues there is torsion, i.e. the movement of the mantle cavity to the front of the animal. Partial de-torsion may occur in later stages of development. IT IS UNRELATED TO SHELL COILING.
What does gastropod torsion achieve?
It improves irrigation but also excretes waste over face.
What are the three taxa within the gastropods?
The Prosobranchiata: gills at the front.
The Opisthobranchiata: gills to the rear after de-torsion.
The Pulmonata: gills lost and mantle cavity functions as simple lung for life in air. Mainly freshwater and terrestrial.
What are the Prosobranchiata like?
A very diverse group of mostly marine spp. Mantle cavities at the front. Shell provides protection against predators. They start small and add tissue to front of shell as they grow.
What is the operculum?
A piece of tissue that can be used to shut of the aperture for complete protection in the case of a predator.
What are the Opisthobranchiata?
Sea slugs and sea hares. Majority have lost their shell. Some have external gills but with unpleasant tasting toxins. Some have cerata containing nemocytes.
What are the Pulmonata?
Terrestrial and freshwater. Snails and slugs. Mantle cavity on the side of the slug.
What are the Bivalvia?
Typically a bivalve shell is two calcareous valves enclosing a visceral mass. The soft tissues are kept in the pallial sinus. Usually sedentary, but exceptions are scallop swimming escape response and Lasaea rubra, bivalve that glides through lichen ‘turf’ in the supralittoral.