Lecture 4: Sphenopsids and ferns: unlikely relations Flashcards
Sphenopsids: living genus example
Equisetum (extant)
Equisetum stem description
has a jointed stem with regular alternations of microphylls in the successive whorles (leaves at the nodes)
In all sphenopsids what does the sporophyte consist of?
a monopodial branch system (one branch)
Sphenopsid: Equisetum leaves
are microphyllous, unbranched and reduced (except in some early fossil forms)
Microphyllous:
one single, unbranched leaf vein
In Equisetum a perennial underground rhizome gives rise to
numerous shoots (root like structure which has a life cycle longer than two years)
Equisetum small plant height
usually 2-50cm in height. (But one species is up to 10m)
Sphenopsid Equisetum: Stomata
are confined to the valleys of the internodes
tracheid:
(an elongated, tapering xylem cell having lignified, pitted, intact walls, adapted for conduction and support)
What type of vascular tissues are present?
tracheids & phloem
Sporangiophores:
(in a plant or fungus) a structure or stalk that bears one or more sporangia
Sphenopsid Equisetum: sporangiophores:
are pelate (having the stalk or support attached to the lower surface at a distance from the margin, as a leaf; shield-shaped) and aggregated (in a cluster) into a terminal strobilus (cone-like structure)
Sphenopsid Equisetum: living homo/heterosporous?
HOMOSPOROUS
some extinct were heterosporous
Sphenopsid Equisetum: living isospores
(one of the spores produced) are rather strange and have pseudoelaters
Sphenopsid Equisetum: microporous or megasporous?
Now all microporous but evidence from fossils of megasporous
Sphenopsid Equisetum: Gametophytes do one of two things
- remain small and produce only antheridia (with a few going on to produce archegonia also)
- become larger and long lived producing archegonia, and if none of these are fertilised also antheridia, followed by a final crop of archegonia. (Encourage outbreeding.)
Sphenopsid Equisetum: Spermatozoids are
multiflagellate
Sphenopsid Equisetum: Embryogeny is
Exoscopic. (Embryogeny in which, of the two cells resulting from the first division of the egg cell, the outer gives rise to the embryo and the inner to the foot. It is seen in the Bryophyta, Psilopsida, and Sphenopsida).